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Folivora
Sloths are a Neotropical realm, Neotropical group of xenarthran mammals constituting the suborder Folivora, including the extant Arboreal locomotion, arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America. Sloths are considered to be most closely related to anteaters, together making up the xenarthran order Pilosa. There are six extant sloth species in two genera – ''Bradypus'' (three-toed sloths) and ''Choloepus'' (two-toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb – although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via parallel evolution from terrestrial ancestors. Besides the extant species, many species of ground sl ...
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Phyllophaga (mammal)
Sloths are a Neotropical group of xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America. Sloths are considered to be most closely related to anteaters, together making up the xenarthran order Pilosa. There are six extant sloth species in two genera – '' Bradypus'' (three-toed sloths) and '' Choloepus'' (two-toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb – although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via parallel evolution from terrestrial ancestors. Besides the extant species, many species of ground sloths ranging up to the size of elephan ...
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Ground Sloth
Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. They varied widely in size with the largest, belonging to genera '' Lestodon'', ''Eremotherium'' and ''Megatherium'', being around the size of elephants. Ground sloths represent a paraphyletic group, as living tree sloths are thought to have evolved from ground sloth ancestors. The early evolution of ground sloths took place during the late Paleogene and Neogene of South America, while the continent was isolated. At their earliest appearance in the fossil record, they were already distinct at the family level. Sloths dispersed into the Greater Antilles during the Oligocene, and the presence of intervening islands between the American continents in the Miocene allowed a dispersal of some species into North America. They were hardy as evidenced by their high species diversity and their presence in a wide variety of environments, extending from the far south of Patagonia ( Cueva del Milodón ...
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Xenarthra
Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a superorder and major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 million years ago, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene. Characteristics Xenarthrans share several characteristics that are not present in other placental mammals, which suggest that xenarthrans desce ...
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Pilosa
The Order (biology), order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes anteaters and sloths (which include the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The Biogeography, biogeographic origins of the Pilosa are still unclear, but they can be traced back in South America as far as the early Paleogene (about 60 million years ago, only a short time after the end of the Mesozoic Era). The presence of these animals in Central America and their former presence in North America is a result of the Great American Interchange. Pilosans of the Caribbean, A number of sloths were also formerly present on the Greater Antilles, Antilles, which they reached from South America by some combination of oceanic dispersal, rafting or floating with the prevailing currents. Together with the armadillos, which are in the order Cingulata, pilosans are part of the larger superorder Xenarthra, a defining charact ...
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Nothrotheriidae
Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately . Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built '' Nothrotheriops'', which reached a length of about . While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today. Evolution During the late Miocene and Pliocene, the sloth genus ''Thalassocnus'' of the west coast of South America became adapted to a shallow-water marine lifestyle. However, the family placement of ''Thalassocnus'' has been disputed; while long considered a nothrotheriid, one 2017 analysis moves it to M ...
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Scelidotheriidae
Scelidotheriidae is a family of extinct ground sloths within the order Pilosa, suborder Folivora and superfamily Mylodontoidea, related to the other extinct mylodontoid family, Mylodontidae, as well as to the living two-toed sloth family Choloepodidae. The only other extant family of the suborder Folivora is the distantly related Bradypodidae. Erected as the family Scelidotheriidae by Florentino Ameghino, Ameghino in 1889, the taxon was demoted to a subfamily of Mylodontidae by Gaudin in 1995. However, recent collagen sequence data indicates the group is less closely related to ''Mylodon'' and ''Lestodon'' than ''Choloepus'' is, and thus it has been elevated back to full family status by Presslee ''et al.'' (2019). Taxonomy Together with Mylodontidae, and the two-toed sloths, the scelidotheriids form the Taxonomic rank, superfamily Mylodontoidea. ''Chubutherium'' is an ancestral and very plesiomorphic member of this family and does not belong to the main group of closely related ...
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Megalocnidae
Megalocnidae is an extinct family (biology), family (alternatively considered to be a superfamily as Megalocnoidea) of sloths, native to the islands of the Greater Antilles from the Early Oligocene to the Mid-Holocene. They are known from Cuba, Hispaniola and Puerto Rico, but are absent from Jamaica. While they were formerly placed in the Megalonychidae alongside two-toed sloths and ground sloths like ''Megalonyx'', recent mitochondrial DNA and collagen sequencing studies place them as the earliest diverging group basal (phylogenetics), basal to all other sloths. or as an outgroup to Megatherioidea. They displayed significant diversity in body size and lifestyle, with ''Megalocnus'' being terrestrial and probably weighing several hundred kilograms, while ''Neocnus'' was likely arboreal and similar in weight to extant tree sloths, at less than 10 kilograms. Origin It is thought that sloths arrived in the Caribbean from South America (where they arose) around the Eocene-Oligocene ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals. They are the only members of the genus ''Bradypus'' (meaning "slow-footed") and the family Bradypodidae. The five living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, the southern maned sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years a ...
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Mylodontidae
Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa, living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another family of extinct ground sloths, Scelidotheriidae, as well as to the extant arboreal two-toed sloths, family Choloepus, Choloepodidae; together these make up the superfamily Mylodontoidea. Phylogenetic analyses based on morphology uncovered the relationship between Mylodontidae and Scelidotheriidae; in fact, the latter was for a time considered a subfamily of mylodontids. However, molecular sequence comparisons were needed for the correct placement of Choloepodidae. These studies have been carried out using mitochondrial DNA sequences as well as with collagen amino acid sequences. The latter results indicate that Choloepodidae is closer to Mylodontidae than Scelidotheriidae is. The only other living sloth family, Bradypodidae (three-toed slot ...
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Megatheriidae
Megatheriidae is a family of Extinction, extinct ground sloths that lived from approximately 23 Annum, mya—11,000 years ago. Megatheriids appeared during the Oligocene, Late Oligocene (Deseadan in the South American land mammal age, SALMA classification), some 29 million years ago, in South America. The group includes the largest known ground sloths, the elephant sized ''Megatherium'' (given its name 'great beast' by Georges Cuvier) and ''Eremotherium''. An early genus that was originally considered a megatheriid, the more slightly built ''Hapalops'', reached a length of about . The nothrotheres have recently been placed in their own family, Nothrotheriidae. The skeletal structure of these ground sloths indicates that the animals were massive. Their thick bones and even thicker joints (especially those on the hind legs) gave their appendages tremendous power that, combined with their size and fearsome claws, provided a formidable defense against predators. The earliest megat ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals. They are the only members of the genus ''Bradypus'' (meaning "slow-footed") and the family Bradypodidae. The five living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, the southern maned sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years a ...
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Two-toed Sloth
''Choloepus'' is a genus of xenarthran mammals from Central and South America within the monotypic family Choloepodidae, consisting of two-toed sloths, sometimes also called two-fingered sloths. The two species of ''Choloepus'' (which means "lame foot" in Ancient Greek), Linnaeus's two-toed sloth (''Choloepus didactylus'') and Hoffmann's two-toed sloth (''Choloepus hoffmanni''), were formerly believed on the basis of morphological studies to be the only surviving members of the sloth family Megalonychidae, but have now been shown by molecular results to be closest to extinct ground sloths of the family Mylodontidae. Extant species Evolution A study of retrovirus and mitochondrial DNA suggests that ''C. didactylus'' and ''C. hoffmani'' diverged 6 to 7 million years ago. Furthermore, based on cytochrome c oxidase subunit I sequences, a similar divergence date ( years ago) between the two populations of ''C. hofmanni'' separated by the Andes has been reported. Th ...
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