Ejectisomes
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Ejectisomes
The cryptomonads (or cryptophytes) are a Class (biology), superclass of algae, most of which have chloroplast, plastids. They are traditionally considered a Division (taxonomy), division of algae among phycologists, under the name of Cryptophyta. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 1 E-5 m, 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagellum, flagella. Some may exhibit mixotrophy. They are classified as superclass Cryptomonada, which is divided into two classes: heterotrophic Goniomonadea and phototrophic Cryptophyceae. The two groups are united under three shared morphological characteristics: presence of a periplast, ejectisomes with secondary scroll, and mitochondrial cristae with flat tubules. Genetic studies as early as 1994 also supported the hypothesis that ''Goniomonas'' was sister to Cryptophyceae. A study i ...
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Cryptophyceae
The cryptophyceae are a class of algae, most of which have plastids. About 230 species are known, and they are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some exhibit mixotrophy. Characteristics Cryptophytes are distinguished by the presence of characteristic extrusomes called ejectosomes or ejectisomes, which consist of two connected spiral ribbons held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectosomes, visible under the light microscope, are associated with the pocket; smaller ones occur underneath the periplast, the cryptophyte-specific cell surrounding. Except for '' Chilomonas'', which has leucoplasts, cryptophytes have one or ...
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Rhodomonas
''Rhodomonas'' is a genus of cryptomonads. It is characterized by its red colour, the square-shaped plates of its inner periplast, its short furrow ending in a gullet, and a distinctly shaped chloroplast A chloroplast () is a type of membrane-bound organelle, organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. Chloroplasts have a high concentration of chlorophyll pigments which captur ... closely associated with its nucleomorph. Historically, ''Rhodomonas'' was characterized by its red chloroplast alone, but this no longer occurs as its taxonomy has become increasingly based on molecular and cellular data. Currently, there is some debate about the taxonomic validity of ''Rhodomonas'' as a genus and further research is needed to verify its taxonomic status. ''Rhodomonas'' is typically found in marine environments, although freshwater reports exist. It is commonly used as a live feed for various aquaculture species. ...
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Goniomonadida
''Goniomonas'' is a genus of Cryptomonads and contains five species. It is a genus of single-celled eukaryotes, including both freshwater and marine species. It lacks plastids, which is very unusual among all of the Cryptophyte genera. It may reflect one of only a small number of times that the Cryptophytes evolved into freshwater habitats. ''Goniomonas'' seems to have a number of freshwater relatives which have not yet been cultured and named. Etymology ''Goniomonas'' means angled small flagellates, combining ''goni'' and ''monas''. History of Discovery It was established by German biologist Samuel Friedrich Stein in 1878. Morphology This genus contains species that are free-swimming, flattened, biflagellate monads. They are oval in lateral view with an obliquely truncate anterior. A furrow extends along the middle of the anterior margin and for a short distance down the ventral margin and is surrounded by a single lateral row of ejectisomes. Chloroplasts, pyrenoids, ...
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Ejectosome
An ejectosome is a cellular organelle responsible for ejecting their contents from the cell. Two unrelated types of ejectosomes are described in the literature: # Cryptomonads have two types of characteristic extrusomes known as ejectosomes. # Intracellular pathogens, such as ''Mycobacterium tuberculosis'', escape from their host cells using an actin-based structure, also called an ejectosome. Cryptomonad ejectosomes contain two connected spiral ribbon-like structures, held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectisomes, visible under the light microscope, are associated with the pocket; smaller ones occur elsewhere on the cell. Mycobacteria are ejected from host cells through the action of an actin-based ejectosome. This escape mechanism requires a cytoskeleton regulator from the host plus an intact mycobacterial ESX-1 secretion Secr ...
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Periplast
The periplast is one of three types of cell-covering of three classes of algae. The cryptomonads have the periplast covering. The Dinophyceae have a type called the amphiesma, and the ''Euglena'' covering is the pellicle. Structure The periplast is a proteinaceous covering. It can be subdivided into an inner periplast and an outer periplast. Both of these components are variable in their composition. The inner periplast may be formed as a single sheet as in the ''Chilomonas paramecium'', or as multiple plates of varying shape. The outer periplast surrounds the plasma membrane. The inner periplast below the plasma membrane may sometimes as in '' Komma caudata'' have a hexagonal arrangement of superficial periplast plates, and these are made up of sub-units. The superficial plates are lined up exactly with the inner plate and are surrounded by crystalline borders with the occasional rosette scale on the surface of the plates. A number of different periplast arrangements have been ...
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Phycocyanin
Phycocyanin is a pigment-protein complex from the light-harvesting phycobiliprotein family, along with allophycocyanin and phycoerythrin. It is an accessory pigment to chlorophyll. All phycobiliproteins are water-soluble, so they cannot exist within the membrane like carotenoids can. Instead, phycobiliproteins aggregate to form clusters that adhere to the membrane called phycobilisomes. Phycocyanin is a characteristic light blue color, absorbing orange and red light, particularly 620 nm (depending on which specific type it is), and emits fluorescence at about 650 nm (also depending on which type it is). Allophycocyanin absorbs and emits at longer wavelengths than phycocyanin C or phycocyanin R. Phycocyanins are found in cyanobacteria (also called blue-green algae). Phycobiliproteins have fluorescent properties that are used in immunoassay kits. Phycocyanin is from the Greek ''phyco'' meaning “algae” and ''cyanin'' is from the English word “cyan", which convention ...
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Nature (journal)
''Nature'' is a British weekly scientific journal founded and based in London, England. As a multidisciplinary publication, ''Nature'' features Peer review, peer-reviewed research from a variety of academic disciplines, mainly in science and technology. It has core editorial offices across the United States, continental Europe, and Asia under the international scientific publishing company Springer Nature. ''Nature'' was one of the world's most cited scientific journals by the Science Edition of the 2022 ''Journal Citation Reports'' (with an ascribed impact factor of 50.5), making it one of the world's most-read and most prestigious academic journals. , it claimed an online readership of about three million unique readers per month. Founded in the autumn of 1869, ''Nature'' was first circulated by Norman Lockyer and Alexander MacMillan (publisher), Alexander MacMillan as a public forum for scientific innovations. The mid-20th century facilitated an editorial expansion for the j ...
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Red Alga
Red algae, or Rhodophyta (, ; ), make up one of the oldest groups of eukaryotic algae. The Rhodophyta comprises one of the largest Phylum, phyla of algae, containing over 7,000 recognized species within over 900 Genus, genera amidst ongoing taxonomic revisions. The majority of species (6,793) are Florideophyceae, and mostly consist of multicellular, ocean, marine algae, including many notable seaweeds. Red algae are abundant in marine habitats. Approximately 5% of red algae species occur in freshwater environments, with greater concentrations in warmer areas. Except for two coastal cave dwelling species in the asexual class Cyanidiophyceae, no terrestrial species exist, which may be due to an evolutionary bottleneck in which the last common ancestor lost about 25% of its core genes and much of its evolutionary plasticity. Red algae form a distinct group characterized by eukaryotic cells without flagella and centrioles, chloroplasts without external endoplasmic reticulum or unstack ...
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Eukaryote
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of Outline of life forms, life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes. The eukaryotes emerged within the archaeal Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati and its sole phylum Promethearchaeota. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as Flagellated cell, flagellated cells. The leading evolutiona ...
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Nucleomorph
Nucleomorphs are small, vestigial eukaryotic nuclei found between the inner and outer pairs of membranes in certain plastids. They are thought to be vestiges of red and green algal nuclei that were engulfed by a larger eukaryote. Because the nucleomorph lies between two sets of membranes, nucleomorphs support the endosymbiotic theory and are evidence that the plastids containing them are complex plastids. Having two sets of membranes indicate that the plastid, a prokaryote, was engulfed by a eukaryote, an alga, which was then engulfed by another eukaryote, the host cell, making the plastid an example of secondary endosymbiosis. Organisms with known nucleomorphs As of 2007, only two monophyletic groups of organisms are known to contain plastids with a vestigial nucleus or nucleomorph: the cryptomonads of the supergroup Cryptista and the chlorarachniophytes of the supergroup Rhizaria, both of which have examples of sequenced nucleomorph genomes. Studies of the genomic organiza ...
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Cell Nucleus
The cell nucleus (; : nuclei) is a membrane-bound organelle found in eukaryote, eukaryotic cell (biology), cells. Eukaryotic cells usually have a single nucleus, but a few cell types, such as mammalian red blood cells, have #Anucleated_cells, no nuclei, and a few others including osteoclasts have Multinucleate, many. The main structures making up the nucleus are the nuclear envelope, a double membrane that encloses the entire organelle and isolates its contents from the cellular cytoplasm; and the nuclear matrix, a network within the nucleus that adds mechanical support. The cell nucleus contains nearly all of the cell's genome. Nuclear DNA is often organized into multiple chromosomes – long strands of DNA dotted with various proteins, such as histones, that protect and organize the DNA. The genes within these chromosomes are Nuclear organization, structured in such a way to promote cell function. The nucleus maintains the integrity of genes and controls the activities of the ...
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