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Dinokaryon
A dinokaryon is a eukaryotic nucleus present in dinoflagellates in which the chromosomes are fibrillar in appearance (i.e. with unmasked DNA fibrils) and are more or less continuously condensed. The nuclear envelope does not break down during mitosis, which is thus termed closed mitosis Mitosis () is a part of the cell cycle in eukaryote, eukaryotic cells in which replicated chromosomes are separated into two new Cell nucleus, nuclei. Cell division by mitosis is an equational division which gives rise to genetically identic ..., or "dinomitosis". The mitotic spindle is extranuclear. Histones are absent. However, recent EST sequencing has revealed the presence of histones in one of the closest relative to dinoflagellates, '' Perkinsus marinus'' and an early-branching dinoflagellate, '' Hematodinium'' sp. However, histone-like proteins of bacterial origins are found in the coding regions at periphery of the dinokaryon's chromosomes. References Dinoflagellate bio ...
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Dinoflagellate
The Dinoflagellates (), also called Dinophytes, are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered protists. Dinoflagellates are mostly marine plankton, but they are also common in freshwater habitats. Their populations vary with sea surface temperature, salinity, and depth. Many dinoflagellates are photosynthetic, but a large fraction of these are in fact mixotrophic, combining photosynthesis with ingestion of prey ( phagotrophy and myzocytosis). In terms of number of species, dinoflagellates are one of the largest groups of marine eukaryotes, although substantially smaller than diatoms. Some species are endosymbionts of marine animals and play an important part in the biology of coral reefs. Other dinoflagellates are unpigmented predators on other protozoa, and a few forms are parasitic (for example, '' Oodinium'' and '' Pfiesteria''). Some dinoflagellates produce resting stages, called dinoflagellate cys ...
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Hematodinium
''Hematodinium'' is a genus of dinoflagellates. Species in this genus, such as ''Hematodinium perezi'', the type species, are internal parasites of the hemolymph of crustaceans such as the Atlantic blue crab (''Callinectes sapidus'') and Norway lobster (''Nephrops norvegicus'').} Species in the genus are economically damaging to commercial crab fisheries, including causing bitter crab disease in the large Tanner or snow crab fisheries of the Bering Sea. Introduction ''Hematodinium'' is a harmful parasitic dinoflagellate in marine decapod crustaceans, often dwelling in the hemolymph of host organisms. Drastic changes in the host's vital organs, tissues, and hemolymph lead to the degeneration of the organism. Previous studies indicate that species in the genus ''Hematodium'' affect many organisms in Order Decapoda, and may also affect Order Amphipoda, and subclass Copepoda. These crustacean taxa include crabs, lobsters, crayfish, prawns, and shrimp – all commercially importan ...
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Eukaryote
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of Outline of life forms, life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes. The eukaryotes emerged within the archaeal Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati and its sole phylum Promethearchaeota. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as Flagellated cell, flagellated cells. The leading evolutiona ...
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Cell Nucleus
The cell nucleus (; : nuclei) is a membrane-bound organelle found in eukaryote, eukaryotic cell (biology), cells. Eukaryotic cells usually have a single nucleus, but a few cell types, such as mammalian red blood cells, have #Anucleated_cells, no nuclei, and a few others including osteoclasts have Multinucleate, many. The main structures making up the nucleus are the nuclear envelope, a double membrane that encloses the entire organelle and isolates its contents from the cellular cytoplasm; and the nuclear matrix, a network within the nucleus that adds mechanical support. The cell nucleus contains nearly all of the cell's genome. Nuclear DNA is often organized into multiple chromosomes – long strands of DNA dotted with various proteins, such as histones, that protect and organize the DNA. The genes within these chromosomes are Nuclear organization, structured in such a way to promote cell function. The nucleus maintains the integrity of genes and controls the activities of the ...
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Chromosome
A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most important of these proteins are the histones. Aided by chaperone proteins, the histones bind to and condense the DNA molecule to maintain its integrity. These eukaryotic chromosomes display a complex three-dimensional structure that has a significant role in transcriptional regulation. Normally, chromosomes are visible under a light microscope only during the metaphase of cell division, where all chromosomes are aligned in the center of the cell in their condensed form. Before this stage occurs, each chromosome is duplicated ( S phase), and the two copies are joined by a centromere—resulting in either an X-shaped structure if the centromere is located equatorially, or a two-armed structure if the centromere is located distally; the jo ...
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Oxyrrhis Marina
''Oxyrrhis marina'' is a species of heterotrophic dinoflagellate with flagella that is widely distributed in the world's oceans. Description This protozoan species has an asymmetrical oval shape to its single-celled body.Lowe, C. D., et al. (2011)Who is ''Oxyrrhis marina''? Morphological and phylogenetic studies on an unusual dinoflagellate.''Journal of Plankton Research'' 33(4) 555-67. It has been likened to a rugby ball.Lowe, C. D., et al. (2011)Collection, isolation and culturing strategies for ''Oxyrrhis marina''.''Journal of Plankton Research'' 33(4) 569-78. The cell usually measures between 20 and 30 micrometers, but it is known to reach 60. It has two flagella with a protruding, tentacle-like bulge between them. The flagella are covered in scales. Most individuals have scales on the body surface, as well. The two flagella have separate functions. One undulates in waves and the other is coiled, producing a corkscrew-like propulsion to move the cell. The individual appears co ...
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Nuclear Envelope
The nuclear envelope, also known as the nuclear membrane, is made up of two lipid bilayer membranes that in eukaryotic cells surround the nucleus, which encloses the genetic material. The nuclear envelope consists of two lipid bilayer membranes: an inner nuclear membrane and an outer nuclear membrane. The space between the membranes is called the perinuclear space. It is usually about 10–50  nm wide. The outer nuclear membrane is continuous with the endoplasmic reticulum membrane. The nuclear envelope has many nuclear pores that allow materials to move between the cytosol and the nucleus. Intermediate filament proteins called lamins form a structure called the nuclear lamina on the inner aspect of the inner nuclear membrane and give structural support to the nucleus. Structure The nuclear envelope is made up of two lipid bilayer membranes, an inner nuclear membrane and an outer nuclear membrane. These membranes are connected to each other by nuclear pores. Two sets ...
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Mitosis
Mitosis () is a part of the cell cycle in eukaryote, eukaryotic cells in which replicated chromosomes are separated into two new Cell nucleus, nuclei. Cell division by mitosis is an equational division which gives rise to genetically identical cells in which the total number of chromosomes is maintained. Mitosis is preceded by the S phase of interphase (during which DNA replication occurs) and is followed by telophase and cytokinesis, which divide the cytoplasm, organelles, and cell membrane of one cell into two new cell (biology), cells containing roughly equal shares of these cellular components. The different stages of mitosis altogether define the mitotic phase (M phase) of a cell cycle—the cell division, division of the mother cell into two daughter cells genetically identical to each other. The process of mitosis is divided into stages corresponding to the completion of one set of activities and the start of the next. These stages are preprophase (specific to plant ce ...
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Mitotic Spindle
In cell biology, the spindle apparatus is the cytoskeletal structure of eukaryotic cells that forms during cell division to separate sister chromatids between daughter cells. It is referred to as the mitotic spindle during mitosis, a process that produces genetically identical daughter cells, or the meiotic spindle during meiosis, a process that produces gametes with half the number of chromosomes of the parent cell. Besides chromosomes, the spindle apparatus is composed of hundreds of proteins. Microtubules comprise the most abundant components of the machinery. Spindle structure Attachment of microtubules to chromosomes is mediated by kinetochores, which actively monitor spindle formation and prevent premature anaphase onset. Microtubule polymerization and depolymerization dynamic drive chromosome congression. Depolymerization of microtubules generates tension at kinetochores; bipolar attachment of sister kinetochores to microtubules emanating from opposite cell poles ...
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Histones
In biology, histones are highly Base (chemistry), basic proteins abundant in lysine and arginine residues that are found in eukaryotic cell nuclei and in most Archaea, Archaeal Phylum, phyla. They act as spools around which DNA winds to create structural units called nucleosomes. Nucleosomes in turn are wrapped into 30-nanometer fibers that form tightly packed chromatin. Histones prevent DNA from becoming tangled and protect it from DNA damage. In addition, histones play important roles in gene regulation and DNA replication. Without histones, unwound DNA in chromosomes would be very long. For example, each human cell has about 1.8 meters of DNA if completely stretched out; however, when wound about histones, this length is reduced to about 9 micrometers (0.09 mm) of 30 nm diameter chromatin fibers. There are five families of histones, which are designated H1/H5 (linker histones), H2, H3, and H4 (core histones). The nucleosome core is formed of two H2A-H2B protein dimer ...
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Perkinsus Marinus
''Perkinsus marinus'' is a species of alveolate belonging to the phylum Perkinsozoa.Joseph, S., et al. (2010)The alveolate ''Perkinsus marinus'': biological insights from EST gene discovery.''BMC Genomics'' 11(1), 228. It is similar to a dinoflagellate. It is known as a prevalent pathogen of oysters, causing massive mortality in oyster populations. The disease it causes is known as dermo or perkinsosis, and is characterized by the degradation of oyster tissues.Bower, S. MSynopsis of infectious diseases and parasites of commercially exploited shellfish: ''Perkinsus marinus'' ("dermo" disease) of oysters.Fisheries and Oceans Canada. 2011. The genome of this species has been sequenced. The species originally was named '' Dermocystidium marinum'' by Mackin, Owen and Collier in 1950. Taxonomy ''P. marinus'' is a protozoan of the protist superphylum Alveolata, the alveolates. Its phylum, Perkinsozoa, is a relatively new taxon positioned between the dinoflagellates and the Apicomple ...
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