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Conchiolin
Conchiolins (sometimes referred to as conchins) are complex proteins which are secreted by a mollusc's outer epithelium (the mantle). These proteins are part of a matrix of organic macromolecules, mainly proteins and polysaccharides, that assembled together form the microenvironment where crystals nucleate and grow. This organic matrix also holds and binds to the crystals of aragonite which give such shells their stiffness. The ions necessary to form calcium carbonate are also secreted by the mantle, but it is the tailored environment created by the organic matrix which causes aragonite (rather than calcite) crystals to nucleate, in much the same way that collagen nucleates hydroxyapatite crystals. Conchiolin serves as a relatively flexible, crack-deflecting matrix for the mineral aggregate particles; its strength and the strong bonding of perlucin can in some cases (such as in the formation of nacre) give the finished material an impressive level of toughness. As ...
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Mollusc
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastr ...
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Periostracum
The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as ''Allonautilus scrobiculatus''. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion. The periostracum is visible as the outer layer of the shell of many molluscan species from terrestrial, freshwater, and marine habitats, and may be seen in land snails, river mussels, and other kinds of freshwater bivalves, as well as in many kinds of marine shelled molluscs. The word ''periostracum'' means "around the shell", meaning that the periostracum is wrapped around what is usually the more calcareous part of the shell. Technically, the calcareous part of the ...
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Periostracum
The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as ''Allonautilus scrobiculatus''. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion. The periostracum is visible as the outer layer of the shell of many molluscan species from terrestrial, freshwater, and marine habitats, and may be seen in land snails, river mussels, and other kinds of freshwater bivalves, as well as in many kinds of marine shelled molluscs. The word ''periostracum'' means "around the shell", meaning that the periostracum is wrapped around what is usually the more calcareous part of the shell. Technically, the calcareous part of the ...
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Mantle (mollusc)
The mantle (also known by the Latin word pallium meaning mantle, robe or cloak, adjective pallial) is a significant part of the anatomy of molluscs: it is the dorsal body wall which covers the visceral mass and usually protrudes in the form of flaps well beyond the visceral mass itself. In many species of molluscs the epidermis of the mantle secretes calcium carbonate and conchiolin, and creates a shell. In sea slugs there is a progressive loss of the shell and the mantle becomes the dorsal surface of the animal. The words mantle and pallium both originally meant cloak or cape, see mantle (vesture). This anatomical structure in molluscs often resembles a cloak because in many groups the edges of the mantle, usually referred to as the ''mantle margin'', extend far beyond the main part of the body, forming flaps, double-layered structures which have been adapted for many different uses, including for example, the siphon. Mantle cavity The ''mantle cavity'' is a central fe ...
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Mollusc Shell
The mollusc (or molluskOften spelled mollusk shell in the USA; the spelling "mollusc" are preferred by ) shell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, which includes snails, clams, tusk shells, and several other classes. Not all shelled molluscs live in the sea; many live on the land and in freshwater. The ancestral mollusc is thought to have had a shell, but this has subsequently been lost or reduced on some families, such as the squid, octopus, and some smaller groups such as the caudofoveata and solenogastres. Today, over 100,000 living species bear a shell; there is some dispute as to whether these shell-bearing molluscs form a monophyletic group (conchifera) or whether shell-less molluscs are interleaved into their family tree. Malacology, the scientific study of molluscs as living organisms, has a branch devoted to the study of shells, and this is called conchology—although ...
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Corneous
Corneous is a biological and medical term meaning horny, in other words made out of a substance similar to that of horns and hooves in some mammals. The word is generally used to describe natural or pathological anatomical structures made out of a hard layer of protein. In mammals this protein would usually be keratin. The word corneous is also often used to describe the operculum of a snail, a gastropod mollusc. Not all gastropods have opercula, but in the great majority of those that do have one, the operculum is corneous. (However in several genera within a few families including the marine Naticidae and the terrestrial Pomatiidae, the operculum is primarily calcareous, in other words mostly made of calcium carbonate.) Corneous opercula are made out of the protein conchiolin Conchiolins (sometimes referred to as conchins) are complex proteins which are secreted by a mollusc's outer epithelium (the mantle). These proteins are part of a matrix of organic macromolecul ...
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Nacre
Nacre ( , ), also known as mother of pearl, is an organicinorganic composite material produced by some molluscs as an inner shell layer; it is also the material of which pearls are composed. It is strong, resilient, and iridescent. Nacre is found in some of the most ancient lineages of bivalves, gastropods, and cephalopods. However, the inner layer in the great majority of mollusc shells is porcellaneous, not nacreous, and this usually results in a non-iridescent shine, or more rarely in non-nacreous iridescence such as ''flame structure'' as is found in conch pearls. The outer layer of cultured pearls and the inside layer of pearl oyster and freshwater pearl mussel shells are made of nacre. Other mollusc families that have a nacreous inner shell layer include marine gastropods such as the Haliotidae, the Trochidae and the Turbinidae. Physical characteristics Structure and appearance Nacre is composed of hexagonal platelets of aragonite (a form of calcium carb ...
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Sporopollenin
270px, SEM image of pollen grains Sporopollenin is one of the most chemically inert biological polymers. It is a major component of the tough outer (exine) walls of plant spores and pollen grains. It is chemically very stable and is usually well preserved in soils and sediments. The exine layer is often intricately sculptured in species-specific patterns, allowing material recovered from (for example) lake sediments to provide useful information to palynologists about plant and fungal populations in the past. Sporopollenin has found uses in the field of paleoclimatology as well. Sporopollenin is also found in the cell walls of several taxa of green alga, including ''Phycopeltis'' (an ulvophycean) and ''Chlorella''. Spores are dispersed by many different environmental factors, such as wind, water or animals. In suitable conditions, the sporopollenin-rich walls of pollen grains and spores can persist in the fossil record for hundreds of millions of years, since sporopollenin is res ...
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Chitin
Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chitin are produced each year in the biosphere. It is a primary component of cell walls in fungi (especially basidiomycetes and filamentous fungi), the exoskeletons of arthropods such as crustaceans and insects, the radulae, cephalopod beaks and gladii of molluscs and in some nematodes and diatoms. It is also synthesised by at least some fish and lissamphibians. Commercially, chitin is extracted from the shells of crabs, shrimps, shellfishes and lobsters, which are major by-products of the seafood industry. The structure of chitin is comparable to cellulose, forming crystalline nanofibrils or whiskers. It is functionally comparable to the protein keratin. Chitin has proved useful for several medicinal, industrial and biotechnological purpose ...
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Toughness
In materials science and metallurgy, toughness is the ability of a material to absorb energy and plastically deform without fracturing."Toughness"
Brian Larson, editor, 2001–2011, The Collaboration for NDT Education, Iowa State University
Toughness is the strength with which the material opposes rupture. One definition of material toughness is the amount of energy per unit volume that a material can absorb before rupturing. This measure of toughness is different from that used for
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Extracellular Matrix
In biology, the extracellular matrix (ECM), also called intercellular matrix, is a three-dimensional network consisting of extracellular macromolecules and minerals, such as collagen, enzymes, glycoproteins and hydroxyapatite that provide structural and biochemical support to surrounding cells. Because multicellularity evolved independently in different multicellular lineages, the composition of ECM varies between multicellular structures; however, cell adhesion, cell-to-cell communication and differentiation are common functions of the ECM. The animal extracellular matrix includes the interstitial matrix and the basement membrane. Interstitial matrix is present between various animal cells (i.e., in the intercellular spaces). Gels of polysaccharides and fibrous proteins fill the interstitial space and act as a compression buffer against the stress placed on the ECM. Basement membranes are sheet-like depositions of ECM on which various epithelial cells rest. Each type ...
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