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Clubmosses
Lycopodiopsida is a class of vascular plants also known as lycopsids, lycopods, or lycophytes. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms ( Lepidodendrales) formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks from div ...
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Lycopodiales
The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 17 accepted genera and about 500 known species. This family originated about 380 million years ago in the early Devonian, though the diversity within the family has been much more recent. "Wolf foot" is another common name for this family due to the resemblance of either the roots or branch tips to a wolf's paw. Description Members of Lycopodiaceae are not spermatophytes and so do not produce seeds. Instead they produce spores, which are oily and flammable, and are the most economically important aspects of these plants. The spores are of one size (i.e. the plants are isosporous) and are borne on a specialized structure at the apex of a shoot called a strobilus (plural: strobili), which resembles a tiny battle club, from which the common name derives. Members of the family share the common feature of having a microphy ...
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Selaginella
''Selaginella'', also known as spikemosses or lesser clubmosses, is a genus of lycophyte. It is usually treated as the only genus in the family Selaginellaceae, with over 750 known species. This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the " fern allies". The species '' S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species '' Selaginella selaginoides'', which turns out (with the closely related '' Selaginella deflexa'') to be a clade that is sister to all other ''Selaginellas'', so any definitive subdivision of the species into separate genera leaves two taxa in ''Selaginella'', with the hundreds of other species in new or resurrected gen ...
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Selaginellales
''Selaginella'', also known as spikemosses or lesser clubmosses, is a genus of lycophyte. It is usually treated as the only genus in the family Selaginellaceae, with over 750 known species. This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the "fern allies". The species '' S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species '' Selaginella selaginoides'', which turns out (with the closely related '' Selaginella deflexa'') to be a clade that is sister to all other ''Selaginellas'', so any definitive subdivision of the species into separate genera leaves two taxa in ''Selaginella'', with the hundreds of other species in new or resurrected genera ...
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Microphylls And Megaphylls
In plant anatomy and evolution a microphyll (or lycophyll) is a type of plant leaf with one single, unbranched leaf vein. Plants with microphyll leaves occur early in the fossil record, and few such plants exist today. In the classical concept of a microphyll, the leaf vein emerges from the protostele without leaving a leaf gap. Leaf gaps are small areas above the node of some leaves where there is no vascular tissue, as it has all been diverted to the leaf. Megaphylls, in contrast, have multiple veins within the leaf and leaf gaps above them in the stem. Leaf vasculature The clubmosses and horsetails have microphylls, as in all extant species there is only a single vascular trace in each leaf. These leaves are narrow because the width of the blade is limited by the distance water can efficiently diffuse cell-to-cell from the central vascular strand to the margin of the leaf. Despite their name, microphylls are not always small: those of '' Isoëtes'' can reach 25 centimetr ...
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Myco-heterotrophy
Myco-heterotrophy (from Greek , , and ) is a symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food from parasitism upon fungi rather than from photosynthesis. A myco-heterotroph is the parasitic plant partner in this relationship. Myco-heterotrophy is considered a kind of cheating relationship and myco-heterotrophs are sometimes informally referred to as "mycorrhizal cheaters". This relationship is sometimes referred to as mycotrophy, though this term is also used for plants that engage in mutualistic mycorrhizal relationships. Relationship between myco-heterotrophs and host fungi Full (or obligate) myco-heterotrophy exists when a non-photosynthetic plant (a plant largely lacking in chlorophyll or otherwise lacking a functional photosystem) gets all of its food from the fungi that it parasitizes. Partial (or facultative) myco-heterotrophy exists when a plant is capable of photosynthesis, but parasitizes fung ...
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Gametophyte
A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the sexual phase in the life cycle of plants and algae. It develops sex organs that produce gametes, haploid sex cells that participate in fertilization to form a diploid zygote which has a double set of chromosomes. Cell division of the zygote results in a new diploid multicellular organism, the second stage in the life cycle known as the sporophyte. The sporophyte can produce haploid spores by meiosis that on germination produce a new generation of gametophytes. Algae In some multicellular green algae ('' Ulva lactuca'' is one example), red algae and brown algae, sporophytes and gametophytes may be externally indistinguishable (isomorphic). In ''Ulva'', the gametes are isogamous, all of one size, shape and general morphology. Land plant ...
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Isoetes
''Isoetes'', commonly known as the quillworts, is a genus of lycopod. It is the only living genus in the family Isoetaceae and order Isoetales. , there were about 200 recognized species, with a cosmopolitan distribution mostly in aquatic habitats but with the individual species often scarce to rare. Species virtually identical to modern quillworts have existed since the Jurassic epoch, though the timing of the origin of modern ''Isoetes'' is subject to considerable uncertainty. The name of the genus may also be spelled ''Isoëtes''. The diaeresis (two dots over the e) indicates that the o and the e are to be pronounced in two distinct syllables. Including this in print is optional; either spelling (''Isoetes'' or ''Isoëtes'') is correct. Description Quillworts are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams, though several (e.g. '' I. butleri'', '' I. histrix'' and '' I. nuttallii'') grow on wet ground that dries out in the summer. The ...
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Palhinhaea Cernua
''Palhinhaea cernua'', synonym ''Lycopodiella cernua'' and ''Lycopodium cernuum'', is a plant in the family Lycopodiaceae, commonly known as the staghorn clubmoss. The Hawaiian name for the plant is ''wāwaeʻiole'', or "rat's foot". It has a substantial number of scientific synonyms in several genera. The genus '' Palhinhaea'' is accepted in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), but not in other classifications which submerge the genus in '' Lycopodiella''. It is the largest of the clubmosses, having rhizomes up to in length, with leafy uprights up to in height. Distribution ''Palhinhaea cernua'' is a widespread pan-tropical species, found mostly at higher elevations in subtropical mountain climates of tropical Africa, Asia, the Pacific Islands and the Neotropics. In Europe, it is found in the Azores (where it is possibly native) and formerly on Madeira. It has been introduced in continental Portugal (Valongo), Sicily and Malta Malta, offici ...
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Fern
The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissues that conduct water and nutrients, and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaf, leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled Fiddlehead fern, fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae (plant), Polypodiidae) and eusporangiate ferns, the latter group including horsetails, Psilotaceae, whisk ferns, marattioid ferns, and ophioglossoid ferns. The fern crown group, consisting of the leptosporangiates and ...
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Isoetales
Isoetales, sometimes also written Isoëtales, is an order of plants in the class Lycopodiopsida. There are about 140-150 living species, all of which are classified in the genus '' Isoetes'' (quillworts), with a cosmopolitan distribution, but often scarce to rare. Living species are mostly aquatic or semi-aquatic, and are found in clear ponds and slowly moving streams. Each leaf is slender and broadens downward to a swollen base up to 5 mm wide where the leaves attach in clusters to a bulb-like, underground corm characteristic of most quillworts. This swollen base also contains male and female sporangia, protected by a thin, transparent covering ( velum), which is used diagnostically to help identify quillwort species. Quillwort species are very difficult to distinguish by general appearance. The best way to identify them is by examining the megaspores under a microscope. '' Isoetes'' are the only living pteridophytes capable of secondary growth. Fossils Some authors i ...
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Incertae Sedis
or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by (of uncertain family), (of uncertain suborder), (of uncertain order) and similar terms. Examples * The fossil plant ''Paradinandra, Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil ''Gluteus minimus (fossil), Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Boc ...
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Vascular Plant
Vascular plants (), also called tracheophytes (, ) or collectively tracheophyta (; ), are plants that have lignin, lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified Tissue (biology), tissue (the phloem) to conduct products of photosynthesis. The group includes most embryophyte, land plants ( accepted known species) excluding mosses. Vascular plants include the clubmosses, Equisetum, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). They are contrasted with nonvascular plants such as mosses and green algae. Scientific names for the vascular plants group include Tracheophyta, Tracheobionta and Equisetopsida sensu lato, Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "hi ...
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