Auxiliary Cell
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Auxiliary Cell
The auxiliary cell is a spore-like structure that forms within the fungal family Gigasporaceae (order Gigasporales). Auxiliary cells have thin cell walls, (spiny), papillate, knobby or sometimes smooth surfaces, and are formed from hyphae after spore germination before the formation of mycorrhizae, and then on the extraradical hyphae in the soil. They may not be 'cells' in the biological sense of the word, as they are structures found with coenocytic hyphae belonging to members of the phylum (division) Glomeromycota. Mostly they are known from members of the Gigasporaceae The Gigasporaceae are a family of fungi in the order Diversisporales. Species in this family are widespread in distribution, and form arbuscular mycorrhiza An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'') is a type of mycorrhiza in w .... Currently this family contains '' Gigaspora'', '' Scutellospora'' and '' Racocetra'', but there are other generic names that have not been widely accepted (''Denti ...
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Spore
In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the Biological life cycle, life cycles of many plants, algae, fungus, fungi and protozoa. They were thought to have appeared as early as the mid-late Ordovician period as an adaptation of early land plants. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. In some rare cases, a diploid spore is also p ...
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Fungus
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one of the kingdom (biology)#Six kingdoms (1998), traditional eukaryotic kingdoms, along with Animalia, Plantae, and either Protista or Protozoa and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of motility, mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related o ...
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Family (biology)
Family (, : ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community ...
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Gigasporaceae
The Gigasporaceae are a family of fungi in the order Diversisporales. Species in this family are widespread in distribution, and form arbuscular mycorrhiza An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'') is a type of mycorrhiza in which the symbiont fungus (''Arbuscular mycorrhizal fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. Arbuscul ... in roots. A species under Gigasporaceae is ''Gigaspora gigantea''. The spores of ''G. gigantea'', found in specific sand dunes, commence in a healthy state of newly formed spores to dead and blackened in seven months through four identifiable steps: they begin as healthy greenish-yellow spores, turn into yellow with brown spots, then reddish-orange-brown, and ultimately dead. A cause of the symptoms of death in spores are soil organisms such as bacteria, protists, and microfauna. References Diversisporales Fungus families {{Glomeromycota-stub ...
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Cell Wall
A cell wall is a structural layer that surrounds some Cell type, cell types, found immediately outside the cell membrane. It can be tough, flexible, and sometimes rigid. Primarily, it provides the cell with structural support, shape, protection, and functions as a selective barrier. Another vital role of the cell wall is to help the cell withstand osmotic pressure and mechanical stress. While absent in many eukaryotes, including animals, cell walls are prevalent in other organisms such as fungi, algae and plants, and are commonly found in most Prokaryote, prokaryotes, with the exception of Mollicutes, mollicute bacteria. The composition of cell walls varies across taxonomic groups, species, cell type, and the cell cycle. In Embryophyte, land plants, the primary cell wall comprises Polysaccharide, polysaccharides like cellulose, hemicelluloses, and pectin. Often, other Polymer, polymers such as lignin, suberin or cutin are anchored to or embedded in plant cell walls. Algae exhibit ...
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Hyphae
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 μm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membran ...
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Mycorrhizae
A mycorrhiza (; , mycorrhiza, or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, the plant root system and its surroundings. Mycorrhizae play important roles in plant nutrition, soil biology, and soil chemistry. In a mycorrhizal association, the fungus colonizes the host plant's root tissues, either intracellularly as in arbuscular mycorrhizal fungi, or extracellularly as in ectomycorrhizal fungi. The association is normally mutualistic. In particular species, or in particular circumstances, mycorrhizae may have a parasitic association with host plants. Definition A mycorrhiza is a symbiotic association between a green plant and a fungus. The plant makes organic molecules by photosynthesis and supplies them to the fungus in the form of sugars or lipids, while the fungus supplies the plant with water and mineral nutrients, such as phosphorus, taken from the soil. Mycorrhiz ...
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Glomeromycota
Glomeromycota (often referred to as glomeromycetes, as they include only one class, Glomeromycetes) are one of eight currently recognized division (mycology), divisions within the Kingdom (biology), kingdom Fungi, with approximately 230 described species. Members of the Glomeromycota form arbuscular mycorrhizas (AMs) with the thalli of bryophytes and the roots of vascular plant, vascular land plants. Not all species have been shown to form AMs, and one, ''Geosiphon pyriformis'', is known not to do so. Instead, it forms an endocytobiotic association with ''Nostoc'' cyanobacterium, cyanobacteria. The majority of evidence shows that the Glomeromycota are dependent on land plants (''Nostoc'' in the case of ''Geosiphon'') for carbon and energy, but there is recent circumstantial evidence that some species may be able to lead an independent existence. The arbuscular mycorrhizal species are terrestrial and widely distributed in soils worldwide where they form symbioses with the roots of ...
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