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Anomalocaridid
Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa '' Anomalocaris canadensis'', '' Hurdia victoria'', '' Peytoia nathorsti'', '' Titanokorys gainessii, Cambroraster falcatus'' and ''Amplectobelua symbrachiata'', the Ordovician '' Aegirocassis benmoulai'' and the Devonian '' Schinderhannes bartelsi''. Etymology The name Radiodonta (L ...
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Anomalocaris
''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of radiodont, an order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygopsis'' Shale of the Stephen Formation in British Columbia, Canada by Joseph Frederick Whiteaves, with more examples found by Charles Doolittle Walcott in the Burgess Shale unit of the Stephen Formation. Other closely related fossils have been found in the older Emu Bay Shale of Australia, as well as possibly elsewhere. Originally several fossilized parts discovered separately (the mouth, frontal appendages and trunk) were thought to be three separate creatures, a misapprehension corrected by Harry B. Whittington and Derek Briggs in a 1985 journal article. With a body length close to 40 centimetres, ''A. canadensis'' is thought to be one of the earliest examples of an apex predator, though others have been found in older Cambrian lagerstätten deposits. Discovery ''An ...
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Dinocaridida
DinocarididaGreek, "Terrible crabs" – sometimes informally spelt Dinocarida, but the second 'id' is linguistically correct – see is a proposed fossil taxon of basal arthropods that flourished in the Cambrian period with occasional OrdovicianVan Roy, P.; Briggs, D. E. G. (2011). "A giant Ordovician anomalocaridid". Nature 473 (7348): 510–513. doi:10.1038/nature09920. edit and Devonian records. Characterized by a pair of frontal appendages and series of body flaps, the name of Dinocaridids comes from Greek, "deinos" and "caris" ("terrible crab"), referring to the suggested role of some of these members as the largest marine predators of their time. Dinocaridids are occasionally referred to as the 'AOPK group' by some literatures, as the group compose of Radiodonta (''Anomalocaris'' and relatives), Opabiniidae (''Opabinia'' and relatives), ''Pambdelurion'' and '' Kerygmachela.'' It is most likely paraphyletic, with ''Kerygmachela'' and ''Pambdelurion'' more basal than the ...
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Peytoia Nathorsti
''Peytoia'' is a genus of hurdiid radiodont that lived in the Cambrian period, containing two species, ''Peytoia nathorsti'' from the Miaolingian of Canada and '' Peytoia infercambriensis'' from Poland, dating to Cambrian Stage 3. Its two frontal appendages had long bristle-like spines, it had no fan tail, and its short stalked eyes were behind its large head. 108 specimens of ''Peytoia'' are known from the Greater Phyllopod bed, where they comprise 0.21% of the community. ''Peytoia nathorsti'' and its junior synonym ''Laggania cambria'' played a major role in the discovery of the radiodont body plan. Initially interpreted as a jellyfish and a sea cucumber respectively, they were eventually shown to be the mouthparts and body of a single animal, which bore ''Anomalocaris''-like appendages. ''Peytoia infercambriensis'' is the geologically oldest known radiodont species. Classification ''Peytoia'' belongs to the clade Hurdiidae, and is closely related to the contemporary genus ...
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Anomalocarididae
Anomalocarididae (occasionally mis-spelt Anomalocaridae) is an extinct family of Cambrian radiodonts, a group of stem-group arthropods. Around 1990s and early 2010s, Anomalocarididae included all radiodont species, hence the previous equivalent of the common name "anomalocaridid" to the whole Radiodonta. This is no longer the case after the revision done by Vinther et al. 2014, as Anomalocarididae restricted to only ''Anomalocaris'' and, if any, a few of closely-related genera since then. Wu et al. 2021 accepted only ''Anomalocaris'' (excluding ''"A." saron'', ''"A." kunmingensis'' and ''"A." briggsi'') and '' Lenisicaris'' as the member of Anomalocarididae, while '' Paranomalocaris'' is questionably included by some studies as well. Anomalocarididae distinguished from the similar family Amplectobeluidae (e.g. '' Amplectobelua'', '' Lyrarapax'') by the presence of triradial oral cone and non-hypertrophied first distal endite. Compared to Hurdiidae and Tamisiocarididae, this ...
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Schinderhannes (genus)
''Schinderhannes bartelsi'' is a species of hurdiid radiodont (anomalocaridid) known from one specimen from the lower Devonian Hunsrück Slates. Its discovery was astonishing because previously, radiodonts were known only from exceptionally well-preserved fossil beds (Lagerstätten) from the Cambrian, 100 million years earlier. Discovery The single specimen was discovered in the Eschenbach-Bocksberg Quarry in Bundenbach, and is named after the outlaw Schinderhannes who frequented the area. Its specific epithet ''bartelsi'' honours Christoph Bartels, a Hunsrück Slate expert. The specimen is now housed in the Naturhistorisches Museum, Mainz. Morphology ''Schinderhannes'' is about long in full body length (6.8cm long excluding telson). Like other radiodonts, the head bears a pair of spiny frontal appendages, a radially-arranged ventral mouthpart ( oral cone), and a pair of large lateral compound eyes. Detailed morphology of the frontal appendages and oral cone are equivoc ...
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Hurdia
''Hurdia'' is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. As a radiodont like '' Peytoia'' and '' Anomalocaris'', it is part of the ancestral lineage that led to euarthropods. Description ''Hurdia'' was one of the largest organisms in the Cambrian oceans, ''H. victoria'' reached approximately in length, while ''H. triangulata'' reached up to just . Its head bore a pair of rake-like frontal appendages which shovelled food into its pineapple-ring-like mouth (oral cone). Like other hurdiids, ''Hurdia'' bore a large frontal carapace protruding from its head composed of three sclerites: a central component known as the H-element and two lateral components known as P-elements. The function of this organ remains mysterious; it cannot have been protective as there was no underlying soft tissue. Originally, it is estimated that body flaps ran along the sides of the organisms, from which large gills were suspended. However, anato ...
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Arthropods
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is "ladder-lik ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Cucumericrus
''Cucumericrus decoratus'' is a species of putative radiodont known from a few poorly preserved specimens. Only fragments of trunk cuticle and corresponded appendages had been revealed, while important radiodont features such as frontal appendages are unknown in this species. The trunk cuticle possess irregular wrinkles and may had been soft in life. Each of the trunk appendage compose of a dorsal flap-like element and a ventral stubby leg with unknown distal region, structurally comparable to the trunk appendages of gill lobopodians (dorsal flaps and ventral lobopods) and euarthropod biramous appendages (flap-like exopod and limb-like endopod). The legs have been interpreted as somewhere between annulated lobopod The lobopodians, members of the informal group Lobopodia (from the Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as ... legs and segmen ...
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Caryosyntrips
''Caryosyntrips'' ("nutcracker") is an extinct genus of radiodont which known from Canada, United States and Spain during the middle Cambrian. ''Caryosyntrips'' is known only from its 14-segmented frontal appendages, which resemble nutcrackers, recovered from the Burgess Shale Formation, Canada Wheeler Shale and Marjum Formation The Marjum Formation is a Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the prece ..., United States, and Valdemiedes Formation, Spain. It was first named by Allison C. Daley, Graham E. Budd in 2010 and the type species is ''Caryosyntrips serratus''. ''Caryosyntrips'' is thought to have used their appendages in a scissor-like grasping or slicing motion, and were probably durophagous, feeding on hard-shelled organisms. Due to the unusual morphology of the frontal appendages and the limi ...
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Hurdiidae
Hurdiidae is an extinct cosmopolitan family of radiodonts, a group of stem-group arthropods, which lived during the Paleozoic Era. It is the most long-lived radiodont clade, lasting from the Cambrian period to the Devonian period. Description Hurdiidae is characterized by frontal appendages with distal region composed of 5 subequal blade-like endites, alongside the enlarged head carapaces and tetraradial mouthpart ( oral cone). The frontal appendages of hurdiids have a distinctive morphology, with the appendage of most species bearing five equally-sized elongate blade-like ventral spines known as endites. Subsequent podomeres were reduced in size and with only small endites or none. Each podomere bore only a single endite, unlike other radiodonts, in which the endites were paired. In most species, the endites were curved medially, so that the appendages formed a basket-like structure. Some hurdiids had greater numbers of endites, with '' Cordaticaris'' bearing seven endi ...
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Amplectobeluidae
''Amplectobeluidae'' is a clade of Cambrian radiodonts. Definition In 2014, ''Amplectobeluidae'' was defined as the most inclusive clade including '' Amplectobelua symbrachiata'' but not '' Anomalocaris canadensis'', '' Tamisiocaris borealis'', or '' Hurdia victoria''. Description Amplectobeluids could be recognized by frontal appendages with well-developed first distal endite, which forming a pincer-like structure that presumably better suited for a grasping function. Complete body fossils of amplectobeluids are only known by '' Amplectobelua'' and '' Lyrarapax'', both showing combination of characters resembling ''Anomalocaris ''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of radiodont, an order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygopsis'' Shale of the Stephen Fo ...'' (i.e. streamlined body; small head with ovoid sclerites; well-developed swimming flaps; a pair o ...
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