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Queen Mandibular Pheromone
Queen mandibular pheromone, or QMP, is a honey bee pheromone produced by the queen and fed to her attendants who share it with the rest of the colony to give the colony the sense of belonging to the queen. Newly emerged queens produce very little QMP. By the sixth day they are producing enough to attract drones for mating. A laying queen makes twice that amount. Lack of QMP seems to attract robber bees. A study of foraging worker bees has suggested that foraging bees are not attracted to QMP. Chemical composition Chemically, QMP is very diverse, with at least 17 major components and other minor ones. Five of these compounds are: 9-oxo-2-decenoic acid (9ODA), ''cis''- and ''trans''-9-hydroxydec-2-enoic acid (9HDA), methyl ''p''-hydroxybenzoate (HOB) and 4-hydroxy-3-methoxyphenylethanol (HVA). Drones QMP functions as a sex pheromone for drones, attracting males to an unmated queen. 9ODA specifically is known to attract drones over long distances, and its combination with 9HDA ...
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Honey Bee
A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmopolitan distribution of honey bees, introducing multiple subspecies into South America (early 16th century), North America (early 17th century), and Australia (early 19th century). Honey bees are known for their construction of perennial colonial nests from wax, the large size of their colonies, and surplus production and storage of honey, distinguishing their hives as a prized foraging target of many animals, including honey badgers, bears and human hunter-gatherers. Only 8 surviving species of honey bees are recognized, with a total of 43 subspecies, though historically 7 to 11 species are recognized. Honey bees represent only a small fraction of the roughly 20,000 known species of bees. The best-known honey bee is t ...
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Hydrophilic
A hydrophile is a molecule or other molecular entity that is attracted to water molecules and tends to be dissolved by water.Liddell, H.G. & Scott, R. (1940). ''A Greek-English Lexicon'' Oxford: Clarendon Press. In contrast, hydrophobes are not attracted to water and may seem to be repelled by it. Hygroscopics ''are'' attracted to water, but are not dissolved by water. Molecules A hydrophilic molecule or portion of a molecule is one whose interactions with water and other polar substances are more thermodynamically favorable than their interactions with oil or other hydrophobic solvents. They are typically charge-polarized and capable of hydrogen bonding. This makes these molecules soluble not only in water but also in polar solvents. Hydrophilic molecules (and portions of molecules) can be contrasted with hydrophobic molecules (and portions of molecules). In some cases, both hydrophilic and hydrophobic properties occur in a single molecule. An example of these amphiphilic ...
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Nasonov Gland
Nasonov's gland produces a pheromone used in recruitment in worker honeybees. The pheromone can serve the purposes of attracting workers to a settled swarm and draw bees who have lost their way back to the Beehive, hive. It is used to recruit workers to food that lacks a characteristic scent and lead bees to water sources. The gland is located on the dorsal side of the abdomen. Its opening is located at the base of the last tergum, tergite at the tip of the abdomen. The gland was first described in 1882 by the Russian zoologist Nikolai Nasonov , Nikolai Viktorovich Nasonov (February 14, 1855 – February 11, 1939). Nasonov thought that the gland performed perspiration; it was Frederick William Lambert Sladen (May 30, 1876 - 1921) of England who in 1901 first proposed that the gland produced a pheromone.See: * (Snodgrass, 1925), p. 116. * * Republished in condensed form in: * Reprinted in: See also *Nasonov pheromone References * Allaby, Michael. ''The Concise Oxford D ...
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Dufour Glands
Dufour's gland is an abdominal gland of certain insects, part of the anatomy of the ovipositor or sting apparatus in female members of Apocrita. The diversification of Hymenoptera took place in the Cretaceous and the gland may have developed at about this time (200 million years ago) as it is present in all three groups of Apocrita, the wasps, bees and ants. Structure Dufour’s gland was first described by Léon Jean Marie Dufour in 1841. Along with the spermatheca and the poison gland, it develops as an invagination of valves of the sternum. It empties at the base of the ovipositor in ants but into the dorsal vaginal wall in bees and wasps. The gland is lined by a single layer of epithelial cells which secrete substances into the hollow interior. Muscles round the opening of the duct and may help control the outflow. Function Dufour’s gland secretes chemicals, the nature and functions of which differs across various hymenopteran groups. The secretion is often used in communic ...
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Queen Retinue Pheromone
Queen retinue pheromones (QRP) are a type of honey bee pheromones, so-called because one of their behavioral effects is to attract a circle of bees (a “retinue”) around the queen. In older literature, the queen pheromone is called ''mandibular pheromone'' because some of its components were first identified from the mandibular glands of queens. ''Retinue pheromone'' may be more accurate because the chemical mix in the pheromone comes from several glands. The following compounds have been identified as present in the QRP, of which, only coniferyl alcohol is found in the mandibular glands. The combination of the five QMP compounds and the four compounds below help create the retinue attraction of worker bees around their queen. * Methyl oleate * Coniferyl alcohol * Hexadecan-1-ol * Alpha-linolenic acid References External links Semiochemicals of Genus ''Apis'' at pherobase.com Beekeeping Insect pheromones {{bee-stub ...
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Juvenile Hormone
Juvenile hormones (JHs) are a group of acyclic sesquiterpenoids that regulate many aspects of insect physiology. The first discovery of a JH was by Vincent Wigglesworth. JHs regulate development, reproduction, diapause, and polyphenisms. In insects, JH (formerly neotenin) refers to a group of hormones, which ensure growth of the larva, while preventing metamorphosis. Because of their rigid exoskeleton, insects grow in their development by successively shedding their exoskeleton (a process known as molting). Juvenile hormones are secreted by a pair of endocrine glands behind the brain called the corpora allata. JHs are also important for the production of eggs in female insects. JH was isolated in 1965 by Karel Sláma and Carroll Williams and the first molecular structure of a JH was solved in 1967. Most insect species contain only juvenile growth hormone (JH) III. To date JH 0, JH I, and JH II have been identified only in the Lepidoptera (butterflies and moths). The for ...
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Mushroom Bodies
The mushroom bodies or ''corpora pedunculata'' are a pair of structures in the Supraesophageal ganglion, brain of arthropods, including insects and Crustacean, crustaceans, and some annelids (notably the ragworm ''Platynereis dumerilii''). They are known to play a role in olfactory memory, olfactory learning and memory. In most insects, the mushroom bodies and the lateral horn of insect brain, lateral horn are the two higher brain regions that receive olfactory information from the antennal lobe via projection neurons. They were first identified and described by French biologist Félix Dujardin in 1850. Structure Mushroom bodies are usually described as neuropils, i.e., as dense networks of neuronal processes (dendrite and axon terminals) and glia. They get their name from their roughly hemispherical ''calyx'', a protuberance that is joined to the rest of the brain by a central nerve tract or ''peduncle''. Most of our current knowledge of mushroom bodies comes from studies of a f ...
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Castes
A caste is a fixed social group into which an individual is born within a particular system of social stratification: a caste system. Within such a system, individuals are expected to marry exclusively within the same caste (endogamy), follow lifestyles often linked to a particular occupation, hold a ritual status observed within a hierarchy, and interact with others based on cultural notions of exclusion, with certain castes considered as either more pure or more polluted than others. The term "caste" is also applied to morphological groupings in eusocial insects such as ants, bees, and termites. The paradigmatic ethnographic example of caste is the division of India's Hindu society into rigid social groups. Its roots lie in South Asia's ancient history and it still exists; however, the economic significance of the caste system in India seems to be declining as a result of urbanisation and affirmative action programs. A subject of much scholarship by sociologists and an ...
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Olfactory Receptor Neuron
An olfactory receptor neuron (ORN), also called an olfactory sensory neuron (OSN), is a sensory neuron within the olfactory system. Structure Humans have between 10 and 20 million olfactory receptor neurons (ORNs). In vertebrates, ORNs are Bipolar neuron, bipolar neurons with dendrites facing the external surface of the cribriform plate with axons that pass through the cribriform foramina with terminal end at olfactory bulbs. The ORNs are located in the olfactory epithelium in the nasal cavity. The cell bodies of the ORNs are distributed among the wikt:stratified, stratified layers of the olfactory epithelium. Many tiny hair-like non-motile cilia protrude from the olfactory receptor cell's dendrites. The dendrites extend to the olfactory epithelial surface and each ends in a dendritic knob from which around 20 to 35 cilia protrude. The cilia have a length of up to 100 micrometres and with the cilia from other dendrites form a meshwork in the olfactory mucus. The surface of ...
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Lymph
Lymph () is the fluid that flows through the lymphatic system, a system composed of lymph vessels (channels) and intervening lymph nodes whose function, like the venous system, is to return fluid from the tissues to be recirculated. At the origin of the fluid-return process, interstitial fluid—the fluid between the cells in all body tissues—enters the lymph capillaries. This lymphatic fluid is then transported via progressively larger lymphatic vessels through lymph nodes, where substances are removed by tissue lymphocytes and circulating lymphocytes are added to the fluid, before emptying ultimately into the right or the left subclavian vein, where it mixes with central venous blood. Because it is derived from interstitial fluid, with which blood and surrounding cells continually exchange substances, lymph undergoes continual change in composition. It is generally similar to blood plasma, which is the fluid component of blood. Lymph returns proteins and excess interstiti ...
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Pheromone
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavior of the receiving individuals. There are ''alarm signal, alarm pheromones'', ''food trail pheromones'', ''sex pheromones'', and many others that affect behavior or physiology. Pheromones are used by many organisms, from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been particularly well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones. The ecological functions and evolution of pheromones are a major topic of research in the field of chemical ecology. Background The portmanteau word "pheromone" was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek language, Greek () and (). Pheromones are also sometimes classified as ec ...
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Antenna (biology)
An antenna (plural: antennae) is one of a pair of appendages used for Sensory system, sensing in arthropods. Antennae are sometimes referred to as ''feelers''. Antennae are connected to the first one or two Segmentation (biology), segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing tactition, touch, air motion, heat, vibration (sound), and especially insect olfaction, smell or gustation, taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate (biology), substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members i ...
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