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Opiliones Penis
The penis of the Opiliones (harvestmen) is an intromittent organ that is not present in other arachnids. It consists of a long shaft (the truncus) and a terminal capsule called a glans, containing a stylus and ejaculatory duct. It may have from one to three muscles, or none as in the specialized lineage Grassatores, where the penis is operated by haemolymph pressure. In some members of the Cyphophthalmi, the structure has been termed a "spermatopositor". This denomination refers to a superficial similarity (analogy) with vertebrate penises and does not have any phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ... relationship with it. References External linksOpiliones reproductive anatomy Dr. Jefferey Schultz, University of MarylandHarvestman Spider penis, ''Thelbunu ...
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Opiliones
The Opiliones (formerly Phalangida) are an Order (biology), order of arachnids, Common name, colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs (see below). , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant taxon, extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their Phylogenetics, phylogenetic position within the Arachnida is disputed; their closest relatives may be camel spiders (Solifugae ...
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Intromittent Organ
An intromittent organ is any external organ of a male organism that is specialized to deliver sperm during copulation. Intromittent organs are found most often in terrestrial species, as most non-mammalian aquatic species fertilize their eggs externally, although there are exceptions. For many species in the animal kingdom, the male intromittent organ is a hallmark characteristic of internal fertilization. Species with intromittent organs Invertebrates Molluscs Male cephalopods have a specialized arm, the hectocotylus, which is inserted into the female's mantle cavity to deliver a spermatophore during copulation. In some species, the hectocotylus breaks off inside the female's mantle cavity; in others, it can be used repeatedly to copulate with different females. Arachnids In spiders, the intromittent organs are the male pedipalps, even though these are not primarily sexual organs, but serve as indirect mating organs; in the male the pedipalps have hollow, clubbed tips, o ...
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Arachnid
Arachnids are arthropods in the Class (biology), class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, opiliones, harvestmen, Solifugae, camel spiders, Amblypygi, whip spiders and Uropygi, vinegaroons. Adult arachnids have eight Arthropod leg, legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs. Almost all Extant taxon, extant arachnids are terrestrial animal, terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 110,000 named species, of which 51,000 are species of spiders. The term is derived from the Ancient Greek, Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, w ...
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Glans
The glans (, : glandes ; from the Latin word for "acorn") is a vascular structure located at the tip of the penis in male mammals or a homologous genital structure of the clitoris in female mammals. Structure The exterior structure of the glans consists of mucous membrane, which is usually covered by foreskin or clitoral hood in naturally developed genitalia. This covering, called the prepuce, is normally retractable in adulthood unless removed by circumcision. The glans naturally joins with the frenulum of the penis or clitoris, as well as the inner labia in women, and the foreskin in men. In non-technical or sexual discussions, often the word "clitoris" refers to the external glans alone, excluding the clitoral hood, frenulum, and internal body of the clitoris. Similarly, phrases "tip" or "head" of the penis refers to the glans alone. Sex differences in humans In males, the glans is known as the glans penis, while in females the glans is known as the clitoral glans. ...
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Stylus
A stylus is a writing utensil or tool for scribing or marking into softer materials. Different styluses were used to write in cuneiform by pressing into wet clay, and to scribe or carve into a wax tablet. Very hard styluses are also used to Engraving, engrave metal, and the slate and stylus system is used to punch out dots to write in Braille. Styluses are held in the hand and thus are usually a narrow elongated shape, similar to a modern ballpoint pen. Many styluses are heavily curved to be held more easily. The word ''stylus'' is also used to describe Stylus (computing), computer styluses used to assist in navigating or providing more precision when using touchscreens. Etymology ''Stylus'' comes from the Latin —the spelling ''stylus'' arose from an erroneous connection with Greek (), 'pillar'.''Oxford Latin Dictionary'', s.v. "stilus" (2012). The Latin word had several meanings, including "a long, sharply pointed piece of metal; the stem of a plant; a pointed instrume ...
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Grassatores
The Grassatores Kury, 2002 are the most diverse infraorder of the Laniatores. It includes over 3,700 species distributed mainly in the tropicsKury, A.B.(2003). Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). '' Revista Ibérica de Aracnología'', vol. especial monográfico 1: 1-337. They are characterized by the male genitalia without musculature, operated by hydraulic pressure and by the double tarsal claws of posterior legs. Superfamilies * Assamioidea Sørensen, 1886 * Epedanoidea Sørensen, 1886 * Gonyleptoidea Sundevall, 1833 * Phalangodoidea Simon, 1879 * Samooidea Sørensen, 1886 * Sandokanoidea Özdikmen & Kury, 2007 * Zalmoxoidea Sørensen, 1886 Beyond the above, there are several other genera such as '' Metapellobunus'' that are often included in the Infraorder, but due to their uncertain taxonomic affinities some researchers have treated as Grassatores ''incertae sedis or is a term used for a taxonomy (biology), taxonomic ...
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Haemolymph
Hemolymph, or haemolymph, is a fluid, similar to the blood in invertebrates, that circulates in the inside of the arthropod's body, remaining in direct contact with the animal's tissues. It is composed of a fluid plasma in which hemolymph cells called hemocyte (invertebrate immune system cell), hemocytes are dispersed. In addition to hemocytes, the plasma also contains many chemicals. It is the major tissue type of the open circulatory system characteristic of arthropods (for example, arachnids, crustaceans and insects). In addition, some non-arthropods such as mollusks possess a hemolymphatic circulatory system. Oxygen-transport systems were long thought unnecessary in insects, but ancestral and functional hemocyanin has been found in the hemolymph. Insect "blood" generally does not carry hemoglobin, although hemoglobin may be present in the tracheal system instead and play some role in respiration. Method of transport In the grasshopper, the closed portion of the system ...
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Cyphophthalmi
Cyphophthalmi is a suborder of harvestmen, colloquially known as mite harvestmen. Cyphophthalmi comprises 36 genera, and more than two hundred described species. The six families are currently grouped into three infraorders: the Boreophthalmi, Scopulophthalmi, and Sternophthalmi. Description Cyphophthalmi are smaller than the more familiar "daddy long-legs" harvestmen, with adults ranging from 1 to 7 mm in length. Moreover, their legs are relatively short compared to most other harvestmen, typically shorter than the body. Some superficially resemble mites, which is where they get their common name. Their coloration is almost always some shade of brown, with a heavily sclerotized body, and they are quite inconspicuous, residing in leaf litter or in caves. Many Cyphophthalmi are eyeless, and presumably rely on olfactory cues to find food and mates. Very little is known about their behavior, though they likely subsist mostly by scavenging and preying on minute arthropods. They ...
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Analogy
Analogy is a comparison or correspondence between two things (or two groups of things) because of a third element that they are considered to share. In logic, it is an inference or an argument from one particular to another particular, as opposed to deduction, induction, and abduction. It is also used where at least one of the premises, or the conclusion, is general rather than particular in nature. It has the general form ''A is to B as C is to D''. In a broader sense, analogical reasoning is a cognitive process of transferring some information or meaning of a particular subject (the analog, or source) onto another (the target); and also the linguistic expression corresponding to such a process. The term analogy can also refer to the relation between the source and the target themselves, which is often (though not always) a similarity, as in the biological notion of analogy. Analogy plays a significant role in human thought processes. It has been argued that analogy li ...
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Vertebrate
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebrata with some 65,000 species, by far the largest ranked grouping in the phylum Chordata. The vertebrates include mammals, birds, amphibians, and various classes of fish and reptiles. The fish include the jawless Agnatha, and the jawed Gnathostomata. The jawed fish include both the Chondrichthyes, cartilaginous fish and the Osteichthyes, bony fish. Bony fish include the Sarcopterygii, lobe-finned fish, which gave rise to the tetrapods, the animals with four limbs. Despite their success, vertebrates still only make up less than five percent of all described animal species. The first vertebrates appeared in the Cambrian explosion some 518 million years ago. Jawed vertebrates evolved in the Ordovician, followed by bony fishes in the Devonian. T ...
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Penis
A penis (; : penises or penes) is a sex organ through which male and hermaphrodite animals expel semen during copulation (zoology), copulation, and through which male placental mammals and marsupials also Urination, urinate. The term ''penis'' applies to many intromittent organs of vertebrates and invertebrates, but not to all. As an example, the intromittent organ of most Cephalopoda is the hectocotylus, a specialized arm, and male spiders use their pedipalps. Even within the Vertebrata, there are morphological variants with specific terminology, such as Hemipenis, hemipenes. Etymology The word "penis" is taken from the Latin word for "Latin profanity#Synonyms and metaphors, tail". Some derive that from Proto-Indo-European language, Indo-European ''*pesnis'', and the Greek word πέος = "penis" from Indo-European ''*pesos''. Prior to the adoption of the Latin word in English, the penis was referred to as a "yard". The Oxford English Dictionary cites an example of the w ...
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