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Melanohalea Exasperatula
''Melanohalea exasperatula'', commonly known as the lustrous camouflage lichen or lustrous brown-shield, is a species of corticolous (bark-dwelling) foliose lichen in the family Parmeliaceae. It has a widespread global distribution and is common in both Europe and northern North America. Its thallus can grow up to in diameter, with marginal up to broad. The upper surface is pale olive-green to red-brown, with isidia (propagules of vegetative reproduction) that are unbranched, inflated, and hollow. It can be distinguished from similar species by the shape and structure of these isidia. The lower surface of the thallus is pale tan to pale brown with scattered, pale rhizines. Apothecia (sexual fruiting bodies) are uncommon, while pycnidia (asexual fruiting bodies) and secondary metabolites have not been observed in this species. The lack of defensive chemicals makes it vulnerable to grazing by slugs and snails. The evolutionary history of ''Melanohalea exasperatula'' is linked t ...
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Helge Thorsten Lumbsch
Helge Thorsten Lumbsch (born 1964) is a German-born lichenologist living in the United States. His research interests include the phylogeny, taxonomy, and phylogeography of lichen-forming fungi; lichen diversity; lichen chemistry and chemotaxonomy. He is the Associate Curator and Head of Cryptogams and Chair of the Department of Botany at the Field Museum of Natural History. Biography Lumbsch was born in Frankfurt in 1964. Interested in lichens already as a schoolboy, he studied natural sciences at the University of Marburg, under the tutelage of Aino Henssen. He received his diploma in 1989, with a dissertation titled ''Ontogenetisch-systematische Studien der Trapeliaceae und verwandter Familien (Lichenisierte Ascomyceten)'' ("Ontogenic-systematic studies of the Trapeliaceae and related families (lichenized ascomycetes"). After Henssen's retirement in 1990, he transferred to the University in Essen, where he worked on the ''Lecanora subfusca'' group in Australasia, a subje ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the la ...
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Melanohalea
''Melanohalea'' is a genus of foliose lichens in the family Parmeliaceae. It contains 30 mostly Northern Hemisphere species that grow on bark or on wood. The genus is characterized by the presence of pseudocyphellae, usually on warts or on the tips of isidia, a non-pored and a medulla containing depsidones or lacking secondary compounds. ''Melanohalea'' was circumscribed in 2004 as a segregate of the morphologically similar genus ''Melanelia''. Taxonomy ''Melanohalea'' was circumscribed in 2004 by lichenologists Oscar Blanco, Ana Crespo, Pradeep K. Divakar, Theodore Esslinger, David L. Hawksworth and H. Thorsten Lumbsch. It is a segregate of ''Melanelia'', a genus created in 1978 to contain the brown '' Parmelia'' species. The circumscription of this genus was questioned later, especially after early molecular phylogenetics studies demonstrated that it was not monophyletic. Subsequently, two genera, '' Melanelixia'' and ''Melanohalea'', were created. ''Melanohalea'' or ...
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Melanelixia
''Melanelixia'' is a genus of foliose lichens in the family Parmeliaceae. It contains 15 Northern Hemisphere species that grow on bark or on wood. The genus is characterized by a pored or fenestrate epicortex (a thin homogeneous polysaccharide layer on the surface of the cortex), and the production of lecanoric acid as the primary chemical constituent of the medulla. ''Melanelixia'' was circumscribed in 2004 as a segregate of the related genus ''Melanelia''. Taxonomy ''Melanelixia'' was circumscribed in 2004 as a segregate of the genus ''Melanelia''. ''Melanelixia'' originally contained eight species, including the type, '' M. glabra''. The genus name combines ''Melanelia'' with the name of lichenologist John A. Elix, "for his immense contributions to lichen systematics and chemistry, especially in Parmeliaceae." Several species distributed in the Southern Hemisphere were removed from ''Melanelixia'' and transferred to the new genus '' Austromelanelixia'' in 2017. This ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have t ...
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Molecular Phylogenetics
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical ...
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Melanelia
''Melanelia'' is a genus of lichenized fungi in the family Parmeliaceae. The genus was circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every po ... by Ted Esslinger in 1978. Species *'' Melanelia commixta'' (Nyl.) A.Thell (1995) *'' Melanelia hepatizon'' (Ach.) A.Thell (1995) *'' Melanelia microglabra'' Divakar, Upreti, G.P.Sinha & Elix (2003) – India *'' Melanelia pseudoglabra'' (Essl.) Essl. (1978) *'' Melanelia stygia'' (L.) Essl. (1978) References Parmeliaceae Lichen genera Lecanorales genera Taxa described in 1978 {{Parmeliaceae-stub ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogene ...
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Hollola
Hollola () is a municipality of Finland, located in the western part of the Päijänne Tavastia region. The municipality is unilingually Finnish and has a population of () and covers an area of of which is water. The population density is . Hollola has the tallest structure in Finland, the Tiirismaa TV-Tower. A popular resort in Hollola is Messilä. There you can do cross-country-skiing, downhill skiing, tramping and play golf. There is a hotel and a camping site in Messilä. Messilä is located by Vesijärvi. In 2018, Hollola hosted the Jukola Relays. This Orienteering event is a night relay, and one of the most popular orienteering events in the world by number of competitors. International relations Twin towns — Sister cities Hollola is twinned with: * Arboga, Sweden * Nordkapp ), North Cape, Norway, other uses, North Cape (other) Nordkapp ( en, North Cape; sme, Davvinjárga or ; fkv, Kappa or ) is a municipality in Troms og Finnmark county, Norw ...
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Johan Petter Norrlin
Johan Petter Norrlin (6 September 1842 in Hollola – 7 January 1917 in Helsinki) was a Finnish botanist and a professor of botany at the University of Helsinki from 1879 to 1903. He was a pioneer of plant geography in Finland, and is also well known for his work on lichens and on the taxonomy of the apomictic taxa of the plant genera '' Hieracium'' and '' Pilosella''. Early life and education Johan Petter Norrlin was born on 6 September 1842 in Hollola, Finland. Norrlin's parents were Nils Nathanael Norrlin and Fredrika Charlotta Lang. He became a student in 1862 at Porvoo high school, and graduated in 1866 as a forester from Evo Forestry College. Norrlin then studied at the University of Helsinki. It was around this time that he met Edvard August Vainio (then named Edvard Lang), the neighbour's son, who was nearly 11 years younger. Young Edvard Lang often accompanied Norrlin on his botanical excursions around Lake Vesijärvi in the summers of 1868 and 1869, helping him coll ...
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Type (biology)
In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set (mathematics), set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the International Code of Nomenclature for algae, fungi, and plants (ICN), the ...
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Parmelia (fungus)
''Parmelia'' is a genus of medium to large foliose lichens.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, It has a global distribution, extending from the ArcticSkult H (1985) A New Subspecies of ''Parmelia omphalodes'' Ascomycetes Described from the Arctic. Annales Botanici Fennici 22, 201-6. to the Antarctic continentD.C. Lindsay (1973) Notes on Antarctic lichens: IV. The genera ''Cetraria'' Hoffm., ''Hypogymnia'' (Nyl.) Nyl., ''Menegazzia'' Massal, ''Parmelia'' Ach. and ''Platismatia'' Culb. et Culb. British Antarctic Survey Bulletin 36, 105-114. but concentrated in temperate regions. There are about 40 species in ''Parmelia''. In recent decades, the once large genus ''Parmelia'' has been divided into a number of smaller genera according to thallus morphology and phylogenetic relatedness. It is a foliaceous lichen, resembling a leaf in shape. The ends of the leaf-like lobes are often squarish-tipped. The upper surface is pale bluish-gra ...
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