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Harpetida
Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period. Harpetid trilobites are characterized among trilobites by bearing a comparatively large, semicircular brim around the cephalon (head) which is often perforated by small pores. This brim is thought to serve as a filter-feeding apparatus. The brim stretches backward on either side of the cephalon (head) and typically has a pronounced suture along the outside. The compound eyes are typically reduced to small tubercles, though they have strong ridges stretching to the glabella (central region of the cephalon). They also typically have 12 or more thoracic segments. The pygidia are usually small. The families of Harpetida were formerly included in the order Ptychopariida, but were recently given their own order (Ebach & McNamara 2002). The subclass Librostoma was erected in 1990 b ...
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Proetida
Proetida is an order of trilobite that lived from the Ordovician to the Permian. It was the last order of trilobite to go extinct, finally dying out in the Permian-Triassic extinction event. Description These typically small trilobites resemble those of the order Ptychopariida, from which the new order Proetida was separated in 1975 by Fortey and Owens. Like the order Phacopida, the proetids have exoskeletons that sometime have pits or small tubercles, especially on the glabella (middle portion of the head). Because of their resemblance to the Ptychopariida in some features, the proetids are included in the subclass Librostoma. Unlike the trilobites of the phacopid suborder Phacopina, whose eyes are schizochroal, the proetids have the more common holochroal eyes. These eyes are characterized by close packing of biconvex lenses beneath a single corneal layer that covers all of the lenses. Each lens is generally hexagonal in outline and in direct contact with the others. ...
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Ptychopariida
Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event. Trilobites have facial sutures that run along the margin of the glabella and/or fixigena to the shoulder point where the cephalon meets the thorax. These sutures outline the cranidium, or the main, central part of the head that does not include the librigena (free cheeks). The eyes are medial along the glabella on the suture line (and some species have no eyes). The fossils of the moults of trilobites can often be told from the fossils of the actual animals by whether the librigena are present. (The librigena, or cheek spines, detach during moulting.) In ptychopariids, short bladelike genal spines are often present on the tips of the librigena. The thorax is large and is typically made up of eight or more segments. ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. Th ...
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Asaphus Lepidurus
''Asaphus'' () is a genus of trilobites that is known from the Lower (upper Arenig) and Middle Ordovician of northwestern Europe (Sweden, Estonia, Saint Petersburg Area). Etymology The generic name is derived from the Greek word ''asaphes'', meaning "indistinct." Description The headshield (or cephalon) and tailshield (or pygidium) are semicircular and without a border (defined by a furrow or a change in convexity parallel to its margin). The cephalon is of approximately equal size as the pygidium (or isopygous). The central raised area of the cephalon (or glabella) is long, reaching the frontal margin. It may have faint lateral glabellar furrows or be smooth, and sometimes an inconspicuous tubercle is present just in front of the hardly discernible occipital ring. The natural fracture lines (sutures) of the head run along the top edges of the compound eye. From the back of the eye these cut to the back of the head (or is said to be opisthoparian) and not to the side. ...
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Richard Fortey
Richard Alan Fortey FRS FRSL (born 15 February 1946 in London) is a British palaeontologist, natural historian, writer and television presenter, who served as president of the Geological Society of London for its bicentennial year of 2007. Early life and education Fortey was educated at Ealing Grammar School for Boys and King's College, Cambridge, where he read Natural Sciences specialising in geology. He received a PhD and DSc from the University of Cambridge. Career Fortey has had a long career as a palaeontologist at the Natural History Museum in London; his research interests include above all, trilobites: at the age of 14, he discovered his first trilobite, sparking a passionate interest that later became a career. He has named numerous trilobite species and still continues his research despite having retired from the Museum. He studies trilobites and graptolites, especially those from the Ordovician and their systematics, evolution and modes of life; he is also involved ...
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Asaphida
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea), but no suborders. Asaphids comprise some 20% of described fossil trilobites. In 2020, the superfamily Trinucleoidea was proposed to be raised to an order ( Trinucleida) and removed from Asaphida. Morphology The Asaphids generally have cephalon (head) and pygidium (tail) parts similar in size, and most species have a prominent median ventral suture. Heads are often flat, and carapace furrows in the head area are often faint or not visible. Thoracic segments typically number 5 - 12, though some species have as few as two and some as many as 30. They also generally have a wide doublure, or rim, that surrounds the cephalon. This causes some specimens to be described as having a charac ...
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Arthropod Subclasses
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is "ladder-lik ...
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Prehistoric Arthropod Taxa
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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