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Juvenile Hormone
Juvenile hormones (JHs) are a group of acyclic sesquiterpenoids that regulate many aspects of insect physiology. The first discovery of a JH was by Vincent Wigglesworth. JHs regulate development, reproduction, diapause, and polyphenisms. In insects, JH (formerly neotenin) refers to a group of hormones, which ensure growth of the larva, while preventing metamorphosis. Because of their rigid exoskeleton, insects grow in their development by successively shedding their exoskeleton (a process known as molting). Juvenile hormones are secreted by a pair of endocrine glands behind the brain called the corpora allata. JHs are also important for the production of eggs in female insects. JH was isolated in 1965 by Karel Sláma and Carroll Williams and the first molecular structure of a JH was solved in 1967. Most insect species contain only juvenile growth hormone (JH) III. To date JH 0, JH I, and JH II have been identified only in the Lepidoptera (butterflies and moths). The for ...
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Sesquiterpenoid
Sesquiterpenes are a class of terpenes that consist of three isoprene units and often have the molecular formula C15H24. Like monoterpenes, sesquiterpenes may be cyclic or contain rings, including many combinations. Biochemical modifications such as oxidation or rearrangement produce the related sesquiterpenoids. It is estimated (2006) that 3000 sesquiterpenes have been identified. Biosynthesis and examples The reaction of geranyl pyrophosphate with isopentenyl pyrophosphate results in the 15-carbon farnesyl pyrophosphate (FPP), which is an intermediate in the biosynthesis of sesquiterpenes such as farnesene. Cyclic sesquiterpenes are more common than cyclic monoterpenes because of the increased chain length and additional double bond in the sesquiterpene precursors. In addition to common six-membered ring systems such as the ones found in zingiberene and bisacurone, cyclization of one end of the chain to the other end can lead to macrocyclic rings such as humulene. Lon ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing more than 150,000 species including horse-flies, crane flies, hoverflies, mosquitoes and others. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great manoeuvrability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the larval food-source and the larvae, which lack true ...
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Ecdysone
Ecdysone is a prohormone of the major insect molting hormone 20-hydroxyecdysone, secreted from the prothoracic glands. It is of steroidal structure. Insect molting hormones (ecdysone and its homologues) are generally called ecdysteroids. Ecdysteroids act as moulting hormones of arthropods but also occur in other related phyla where they can play different roles. In ''Drosophila melanogaster'', an increase in ecdysone concentration induces the expression of genes coding for proteins that the larva requires. It causes chromosome puffs (sites of high expression) to form in polytene chromosomes. Recent findings in the laboratory of Chris Q. Doe have found a novel role of this hormone in regulating temporal gene transitions within neural stem cells of the fruit fly. Ecdysone and other ecdysteroids also appear in many plants mostly as a protection agent ( toxins or antifeedants) against herbivorous insects. These phytoecdysteroids have been reputed to have medicinal value. Th ...
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Diminutive
A diminutive is a word obtained by modifying a root word to convey a slighter degree of its root meaning, either to convey the smallness of the object or quality named, or to convey a sense of intimacy or endearment, and sometimes to belittle something or someone. A ( abbreviated ) is a word-formation device used to express such meanings. A is a diminutive form with two diminutive suffixes rather than one. Purpose Diminutives are often employed as nicknames and pet names when speaking to small children and when expressing extreme tenderness and intimacy to an adult. The opposite of the diminutive form is the augmentative. In some contexts, diminutives are also employed in a pejorative sense to denote that someone or something is weak or childish. For example, one of the last Western Roman emperors was Romulus Augustus, but his name was diminutivized to "Romulus Augustulus" to express his powerlessness. Formation In many languages, diminutives are word forms that ...
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Rhodnius
''Rhodnius'' is a genus of assassin bugs in the subfamily Triatominae (the kissing bugs), and is an important vector in the spread of Chagas disease. The ''Rhodnius'' species were important models for Sir Vincent Wigglesworth's studies of insect physiology, specifically growth and development. Species *''Rhodnius amazonicus'' Almeida, Santos & Sposina, 1973 *''Rhodnius brethesi'' Matta, 1919 (Tc) *''Rhodnius colombiensis'' Moreno Mejía, Galvão & Jurberg, 1999 *''Rhodnius dalessandroi'' Carcavallo & Barreto, 1976 *''Rhodnius domesticus'' Neiva & Pinto, 1923 (Tc) *''Rhodnius ecuadoriensis'' Lent & León, 1958 (Tc) *''Rhodnius milesi'' Carcavallo, Rocha, Galvão, Jurberg, 2001 *'' Rhodnius nasutus'' Stål, 1859 (Tc) *'' Rhodnius neglectus'' Lent, 1954 (Tc) *''Rhodnius neivai'' Lent, 1953 *'' Rhodnius pallescens'' Barber, 1932 (Tc) (principal vector in Panama). *''Rhodnius paraensis'' Sherlock, Guitton & Miles, 1977 (Tc) *''Rhodnius pictipes'' Stål, 1872 (Tc) *'' Rhodnius prolixus ...
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Instar
An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form. Differences between instars can often be seen in altered body proportions, colors, patterns, changes in the number of body segments or head width. After shedding their exoskeleton (moulting), the juvenile arthropods continue in their life cycle until they either pupate or moult again. The instar period of growth is fixed; however, in some insects, like the salvinia stem-borer moth, the number of instars depends on early larval nutrition. Some arthropods can continue to moult after sexual maturity, but the stages between these subsequent moults are generally not called instars. For most insect species, an ''instar'' is the developmental stage of the larval forms of holometabolous (complete metamorphism) or ny ...
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Ecdysis
Ecdysis is the moulting of the cuticle in many invertebrates of the clade Ecdysozoa. Since the cuticle of these animals typically forms a largely inelastic exoskeleton, it is shed during growth and a new, larger covering is formed. The remnants of the old, empty exoskeleton are called exuviae. After moulting, an arthropod is described as ''teneral'', a ''callow''; it is "fresh", pale and soft-bodied. Within one or two hours, the cuticle hardens and darkens following a Tanning (leather), tanning process analogous to the production of leather. During this short phase the animal expands, since growth is otherwise constrained by the rigidity of the exoskeleton. Growth of the limbs and other parts normally covered by the hard exoskeleton is achieved by transfer of body fluids from soft parts before the new skin hardens. A spider with a small abdomen may be undernourished but more probably has recently undergone ecdysis. Some arthropods, especially large insects with tracheal respira ...
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Juvenile Hormone Epoxide Hydrolase
Juvenile hormone epoxide hydrolase (JHEH) is an enzyme that inactivates insect juvenile hormones. This inactivation is accomplished through hydrolysis of the epoxide functional group contained within these hormones into diols. JHEH is one of two enzymes involved in the termination of signaling properties of the various juvenile hormones. The other is juvenile-hormone esterase, or JHE. The first observation of activity from JHEH was in '' Rhodnius prolixus'' and '' Schistocerca gregaria''. JH esterase was also first observed in this same work. Mechanism JH esterase and epoxide hydrolase are the only known enzymes involved in primary degradation of JH. The inactivated carboxylic acid products of juvenile hormone esterase can be reactivated in peripheral tissues by esterification restoring its biological activity. In contrast, the juvenile hormone diols that are the product of the epoxide hydrolase are very hydrophilic and cannot be converted back to JH. Thus, this enzyme perma ...
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Haemolymph
Hemolymph, or haemolymph, is a fluid, similar to the blood in invertebrates, that circulates in the inside of the arthropod's body, remaining in direct contact with the animal's tissues. It is composed of a fluid plasma in which hemolymph cells called hemocyte (invertebrate immune system cell), hemocytes are dispersed. In addition to hemocytes, the plasma also contains many chemicals. It is the major tissue type of the open circulatory system characteristic of arthropods (for example, arachnids, crustaceans and insects). In addition, some non-arthropods such as mollusks possess a hemolymphatic circulatory system. Oxygen-transport systems were long thought unnecessary in insects, but ancestral and functional hemocyanin has been found in the hemolymph. Insect "blood" generally does not carry hemoglobin, although hemoglobin may be present in the tracheal system instead and play some role in respiration. Method of transport In the grasshopper, the closed portion of the system ...
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Allatostatin
Allatostatins are neuropeptide hormones in insects and crustacea. They have a twofold function: they both inhibit the generation of juvenile hormone and reduce their food intake. They are therefore putative targets for insecticide research. Types There are three distinct Allatostatin types: A, B, and C. Allatostatin C's have 3 subtypes as a result of gene multiplication: C, CC, and CCC. Each Allatostatin type has a unique evolutionary history resulting in distinct conservation and functions across the animal kingdom. Although originally identified in different insects, all three type are found in ''Drosophila (needs source)''. Allatostatin A Allatostatin A (AstA) peptides are found in all arthropods and contain a C-terminus Y/FXFGLamide. In Drosophila, there are 4 AstA peptides (AstA-1, AstA-2, AstA-3, AstA-4) and 2 AstA receptors (AstA-R1 and AstA-R2). The AstA receptor is a GIRK1 channel and is homologous to the mammalian galanin receptor{{Cite journal , last1=Birgül , firs ...
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G-protein Coupled Receptor
G protein-coupled receptors (GPCRs), also known as seven-(pass)-transmembrane domain receptors, 7TM receptors, heptahelical receptors, serpentine receptors, and G protein-linked receptors (GPLR), form a large protein family, group of evolutionarily related proteins that are cell surface receptors that detect molecules outside the cell (biology), cell and activate cellular responses. They are coupled with G proteins. They pass through the cell membrane seven times in the form of six loops (three extracellular loops interacting with ligand molecules, three intracellular loops interacting with G proteins, an N-terminus, N-terminal extracellular region and a C-terminal intracellular region) of amino acid residues, which is why they are sometimes referred to as seven-transmembrane receptors. Text was copied from this source, which is available under Attribution 2.5 Generic (CC BY 2.5) licence/ref> Ligands can bind either to the extracellular N-terminus and loops (e.g. glutama ...
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Conifer
Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class (biology), class, Pinopsida. All Neontology, extant conifers are perennial plant, perennial woody plants with secondary growth. The majority are trees, though a few are shrubs. Examples include Cedrus, cedars, Pseudotsuga, Douglas-firs, Cupressaceae, cypresses, firs, junipers, Agathis, kauri, larches, pines, Tsuga, hemlocks, Sequoioideae, redwoods, spruces, and Taxaceae, yews.Campbell, Reece, "Phylum Coniferophyta". ''Biology''. 7th ed. 2005. Print. p. 595. As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species. Although the total number of species is relatively small, conifers are ecology, ecologically important. They are the dominant plants over large areas of land, most notably ...
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