Hoffstetterius
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Hoffstetterius
''Hoffstetterius'' is an extinct genus of toxodontid notoungulate mammal, belonging to the subfamily Toxodontinae whose remains were discovered in the Middle to Late Miocene (Mayoan to Montehermosan) Mauri Formation in the La Paz Department in Bolivia.''Hoffstetterius''
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The only described species is the ''Hoffstetterius imperator''.Saint-Andre, P. A. (1993). ''Hoffstetterius imperator'' ng, n. sp. du Miocène supérieur de l'Altiplano bolivien et le statut des Dinotoxodontinés (Mammalia, Notoungulata). Com ...
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Toxodontinae
Toxodontidae is an extinct family of notoungulate mammals, known from the Oligocene to the Holocene (11,000 BP) of South America, with one genus, '' Mixotoxodon'', also known from the Pleistocene of Central America and southwestern North America (Texas).E. Lundelius, et al. 2013. The first occurrence of a toxodont (Mammalia, Notoungulata) in the United States. ''Journal of Vertebrate Paleontology'', Vol 33, No 1, pp. 229–23DOI:10.1080/02724634.2012.711405/ref> They somewhat resembled rhinoceroses, and had teeth with high crowns and open roots, suggesting that they often fed on tough pampas grass. However, isotopic analyses have led to the conclusion that the most recent forms were grazing and browsing generalists. Taxonomy The endemic notoungulate and litoptern ungulates of South America have been shown by studies of collagen and mitochondrial DNA sequences to be a sister group to the perissodactyl Odd-toed ungulates, mammals which constitute the taxonomic order Per ...
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Toxodontid
Toxodontidae is an extinct family of notoungulate mammals, known from the Oligocene to the Holocene (11,000 BP) of South America, with one genus, '' Mixotoxodon'', also known from the Pleistocene of Central America and southwestern North America (Texas).E. Lundelius, et al. 2013. The first occurrence of a toxodont (Mammalia, Notoungulata) in the United States. ''Journal of Vertebrate Paleontology'', Vol 33, No 1, pp. 229–23DOI:10.1080/02724634.2012.711405/ref> They somewhat resembled rhinoceroses, and had teeth with high crowns and open roots, suggesting that they often fed on tough pampas grass. However, isotopic analyses have led to the conclusion that the most recent forms were grazing and browsing generalists. Taxonomy The endemic notoungulate and litoptern ungulates of South America have been shown by studies of collagen and mitochondrial DNA sequences to be a sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises th ...
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Notoungulate
Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined during the Late Neogene, with only the large toxodontids persisting until the end of the Pleistocene. Collagen analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls (odd-toed ungulates), including rhinoceroses, tapirs and equines. but their relationships to other South American ungulates are uncertain. ...
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Femur
The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with the tibia (shinbone) and patella (kneecap), forming the knee joint. By most measures the two (left and right) femurs are the strongest bones of the body, and in humans, the largest and thickest. Structure The femur is the only bone in the upper leg. The two femurs converge medially toward the knees, where they articulate with the proximal ends of the tibiae. The angle of convergence of the femora is a major factor in determining the femoral-tibial angle. Human females have thicker pelvic bones, causing their femora to converge more than in males. In the condition ''genu valgum'' (knock knee) the femurs converge so much that the knees touch one another. The opposite extreme is ''genu varum'' (bow-leggedness). In the general pop ...
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Postcrania
Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated skeletal elements; these are referred to as ''postcrania''. There is some disagreement over whether the skull and skeleton belong to the same or different animals. One example is the case of a Cretaceous sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ... skull of '' Nemegtosaurus'' found in association with the postcranial skeleton '' Opisthocoelicaudia''. In paleoanthropological studies, reconstruction of relationship between various species/remains is considered to be better supported by cranial characters rather than postcrania ...
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Axis (anatomy)
In anatomy, the axis (from Latin ''axis'', "axle") or epistropheus is the second cervical vertebra (C2) of the spine, immediately inferior to the atlas, upon which the head rests. The axis' defining feature is its strong odontoid process (bony protrusion) known as the dens, which rises dorsally from the rest of the bone. Structure The body is deeper in front or in the back and is prolonged downward anteriorly to overlap the upper and front part of the third vertebra. It presents a median longitudinal ridge in front, separating two lateral depressions for the attachment of the longus colli muscles. Odontoid Process of Axis (Dens) The dens, also called the odontoid process or the peg, is the most pronounced projecting feature of the axis. The dens exhibits a slight constriction where it joins the main body of the vertebra. The condition where the dens is separated from the body of the axis is called ''os odontoideum'' and may cause nerve and circulation compression syndrom ...
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Humerus
The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a rounded head, a narrow neck, and two short processes (tubercles, sometimes called tuberosities). The body is cylindrical in its upper portion, and more prismatic below. The lower extremity consists of 2 epicondyles, 2 processes (trochlea & capitulum), and 3 fossae (radial fossa, coronoid fossa, and olecranon fossa). As well as its true anatomical neck, the constriction below the greater and lesser tubercles of the humerus is referred to as its surgical neck due to its tendency to fracture, thus often becoming the focus of surgeons. Etymology The word "humerus" is derived from la, humerus, umerus meaning upper arm, shoulder, and is linguistically related to Gothic ''ams'' shoulder and Greek ''ōmos''. Structure Upper extremity The upper or pr ...
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Radius
In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the spoke of a chariot wheel. as a function of axial position ../nowiki>" Spherical coordinates In a spherical coordinate system, the radius describes the distance of a point from a fixed origin. Its position if further defined by the polar angle measured between the radial direction and a fixed zenith direction, and the azimuth angle, the angle between the orthogonal projection of the radial direction on a reference plane that passes through the origin and is orthogonal to the zenith, and a fixed reference direction in that plane. See also *Bend radius *Filling radius in Riemannian geometry *Radius of convergence * Radius of convexity *Radius of curvature *Radius of gyration ''Radius of gyration'' or gyradius of a body about the axis of r ...
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Ulna
The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of the forearm as the little finger. It runs parallel to the radius, the other long bone in the forearm. The ulna is usually slightly longer than the radius, but the radius is thicker. Therefore, the radius is considered to be the larger of the two. Structure The ulna is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. It is broader close to the elbow, and narrows as it approaches the wrist. Close to the elbow, the ulna has a bony process, the olecranon process, a hook-like structure that fits into the olecranon fossa of the humerus. This prevents hyperextension and forms a hinge joint with the trochlea of the humerus. There ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the la ...
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Paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Often there is more than one paratype. Paratypes are usually held in museum research collections. The exact meaning of the term ''paratype'' when it is used in zoology is not the same as the meaning when it is used in botany. In both cases however, this term is used in conjunction with ''holotype''. Zoology In zoological nomenclature, a paratype is officially defined as "Each specimen of a type series other than the holotype.", ''International Code of Zoological Nomenclature'' In turn, this definition relies on the definition of a "type series". A type series is the material (specimens of organisms) that was cited in the original publication of the new species or subspecies, and was not excluded from being type material by the author (t ...
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Chasicoan
The Chasicoan ( es, Chasiquense or es, Chasicoense) age is a period of geologic time from 10–9 Ma within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Mayoan and precedes the Huayquerian age.Chasicoan
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Etymology

The Chasicoan is named after the
Arroyo Chasicó Formation Arroyo often refers to: * Arroyo (creek), an intermittently dry creek Arroyo may also refer to: People * Arroyo (surname) Places United St ...
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