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Hesperotestudo Incisa
''Hesperotestudo'' ("Western turtle") is an extinct genus of tortoise native to North and Central America (ranging as far south as Costa Rica) from the Early Miocene to the Late Pleistocene. Species of ''Hesperotestudo'' varied widely in size, with a large undescribed specimen from the Late Pleistocene of El Salvador reaching in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of ''Geochelone'', it is now considered to be distantly related to that genus. Its relationships with other tortoises are uncertain. The exposed areas of the bodies of ''Hesperotestudo'' species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of ''Hesperotestudo'' were relatively tolerant of cold weather. ''Hesperotestudo'' became extinct at the end of the Pleistocene roughly co-incident with the arrival of the first humans in North America. There is apparently a site in Florida wh ...
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Extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and recover. As a species' potential Range (biology), range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxon, Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the Fossil, fossil record) after a period of apparent absence. Over five billion species are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryotes globally, possibly many times more if microorganisms are included. Notable extinct animal species include Dinosaur, non-avian dinosaurs, Machairodontinae, saber-toothed cats, and mammoths. Through evolution, species arise through the process of specia ...
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Langhans Giant Cell
Langhans giant cells (LGC) are giant cells found in granulomatous conditions. They are formed by the fusion of epithelioid cells (macrophages), and contain nuclei arranged in a horseshoe-shaped pattern in the cell periphery. Although traditionally their presence was associated with tuberculosis, they are not specific for tuberculosis or even for mycobacterial disease. In fact, they are found in nearly every form of granulomatous disease, regardless of etiology. Terminology Langhans giant cells are named after Theodor Langhans (1839–1915), a German pathologist. Causes In 2012, a research paper showed that when activated CD4+ T cells and monocytes are in close contact, interaction of CD40-CD40L between these two cells and subsequent IFNγ secretion by the T cells causes upregulation and secretion of fusion-related molecule DC-STAMP (dendritic cell-specific transmembrane protein) by the monocytes, which results in LGC formation. Clinical significance Langhans giant cells ...
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Cenozoic Reptiles Of North America
The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three geological eras of the Phanerozoic Eon, preceded by the Mesozoic and Paleozoic. The Cenozoic started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor. The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammalsthe eutherians (placentals) in the Northern Hemisphere and the metatherians (marsupials, now mainly restricted to Australia and to some extent South America) in the Southern Hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that large mamm ...
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Pleistocene Turtles
The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek () 'most' and (; Latinized as ) 'new'. The aridification and cooling trends of the preceding Neogene were continued in the Pleistocene. The climate was strongly variable depending on the glacial cycle, oscillating between cold Glacial period, glacial periods and warmer Interglacial, int ...
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Pliocene Turtles
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58See the 2014 version of the ICS geologic time scale
million years ago (Ma). It is the second and most recent epoch of the Neogene Period in the Cenozoic, Cenozoic Era. The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in the Pleistocene. As with other older geologic periods, the Stratum, geological strata that define the start and end are well-identified but the exact dates of the start a ...
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Miocene Turtles
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene followed the Oligocene and preceded the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including the dispersal of proboscideans and hominoids into Eurasia. During the late Miocene, the connections between the ...
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Testudinidae
Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backward into the shell to protect them. Tortoises can vary in size with some species, such as the Galápagos giant tortoise, growing to more than in length, whereas others like the Speckled cape tortoise have shells that measure only long. Several lineages of tortoises have independently evolved very large body sizes in excess of , including the Galapagos giant tortoise and the Aldabra giant tortoise. They are usually diurnal animals with tendencies to be crepuscular depending on the ambient temperatures. They are generally reclusive animals. Tortoises are the longest-living land animals in the world, although the longest-living species of tortoise ...
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Burdigalian
The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (stage), Aquitanian, the Burdigalian was the first and longest warming period of the MioceneEdward Petuch, Ph.D. Florida Atlantic University, Department of Geosciences. and is succeeded by the Langhian. Stratigraphic definition The name Burdigalian comes from ''Burdigala'', the Latin name for the city of Bordeaux, France. The Burdigalian Stage was introduced in scientific literature by Charles Depéret in 1892. The base of the Burdigalian is at the first appearance of foram species ''Globigerinoides altiaperturus'' and the top of magnetic chronozone C6An. , an official GSSP for the Burdigalian had not yet been assigned. The top of the Burdigalian (the base of the Langhian) is defined by the first appearance of foram species ''Praeorbulina gl ...
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Oakville Formation
The Oakville Formation is a geologic formation in Texas. It preserves fossils dating back to the Neogene period. See also * List of fossiliferous stratigraphic units in Texas * Paleontology in Texas Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has said that "Texas is as big for fossils as it is for everything else." Some of the most import ... References * Neogene geology of Texas {{Neogene-stub ...
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Zanclean
The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and followed by the Piacenzian Age. The Zanclean can be correlated with regionally used stages, such as the Opoitian of New Zealand, and the Tabianian or Dacian of Central Europe. It also corresponds to the late Hemphillian to mid-Blancan North American Land Mammal Ages. In California, the Zanclean roughly corresponds to the middle part of the Delmontian stage. Definition The Zanclean Stage was introduced by Giuseppe Seguenza in 1868. It is named after '' Zancle'', the pre- Roman name for the Italian city of Messina on Sicily. The base of the Zanclean (and the Pliocene Series) lies with the top of magnetic chronozone Cr3 (about 100,000 years before the Thvera normal subchronozone C3n.4n). The base is also close to the extinction level of th ...
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Tortonian
The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tortonian roughly overlaps with the regional Pannonian Stage of the Paratethys timescale of Central Europe. It also overlaps the upper Astaracian, Vallesian and lower Turolian European land mammal ages, the upper Clarendonian and lower Hemphillian North American land mammal ages and the upper Chasicoan and lower Huayquerian South American land mammal ages. Definition The Tortonian was introduced by Swiss stratigrapher Karl Mayer-Eymar in 1858. It was named after the Italian city of Tortona in the Piedmont region. The base of the Tortonian Stage is at the last common appearance of calcareous nanoplankton '' Discoaster kugleri'' and planktonic foram '' Globigerinoides subquadratus''. It is also associated with the short normal pol ...
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Hemphillian
The Hemphillian North American Stage on the geologic timescale is a North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 10,300,000 to 4,900,000 years BP. It is usually considered to overlap the Tortonian age of the Late Miocene and Zanclean age of the Early Pliocene. The Hemphillian is preceded by the Clarendonian and followed by the Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years BP, a period of .
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