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Graphidales Genera
Graphidales is an order (biology), order of lichen-forming fungi in the class Lecanoromycetes. It contains 6 family (biology), families, about 81 genus, genera and about 2,228 species. Family Graphidaceae are the largest crustose family within Graphidales order comprising more than 2000 species, which are widely distributed in tropical and subtropical regions of the world. History The Graphidales were introduced in a 1884 publication by Frigyes Ákos Hazslinszky in Magyar Birodalom Zuzmó-Flórája on page 216 as family Graphideae. In 1907, they were established as an Order (biology), order by American botanist Charles Edwin Bessey, Bessey (1845–1915), When the order was introduced, it contained just two families, the Graphidaceae and Thelotremataceae who were both mainly tropical based and each family had about 800–1000 species. Sherwood in 1977 proposed to maintain a distinction between the Graphidales with mostly lichenised members and the Ostropales which included mostl ...
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Pallidogramme Chrysenteron
''Pallidogramme'' is a genus of lichen-forming fungi in the family Graphidaceae. It has 8 species of corticolous lichen, corticolous (bark-dwelling), crustose lichens. Taxonomy ''Pallidogramme'' was proposed in 2008 by the lichenologists Bettina Staiger, Klaus Kalb, and Robert Lücking, as a replacement name for ''Leucogramma'' . The justification for this change originated from the previously established ''Hemithecium'' subgen. ''Leucogramma'' by Staiger in 2002. There was, however, a pre-existing generic name, ''Leucogramma'' , which was based on the same species as Staiger's subgenus, ''Hemithecium chrysenteron''. Both of these names were unusable at the genus level due to their Homonym (biology), homonymy with ''Leucogramma'' a name with unknown taxonomy (biology), taxonomic affinity. As a result, a new name was deemed necessary to avoid confusion and maintain nomenclatural clarity. Description The defining features of genus ''Pallidogramme'' include a crustose lichen, cr ...
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Crustose
Crustose is a Habit (biology), habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the Substrate (biology), substrates at all points. ''Crustose'' is found on Rock (geology), rocks and Bark (botany), tree bark. Some species of marine algae of the Rhodophyta, in particular members of the Order (biology), order Corallinales, Family (biology), family Corallinaceae, subfamily Melobesioideae with cell walls containing calcium carbonate grow to great depths in the intertidal zone, forming crusts on various substrates. The substrate can be rocks throughout the intertidal zone, or, as in the case of the Corallinales, reef-building coral reef, corals, and other living organisms including plants, such as mangroves and animals such as shelled molluscs. The coralline red algae are major members of coral reef communities, cementing the corals together with their crusts. Among the ...
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Green Algae
The green algae (: green alga) are a group of chlorophyll-containing autotrophic eukaryotes consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ( Embryophytes) have emerged deep within the charophytes as a sister of the Zygnematophyceae. Since the realization that the Embryophytes emerged within the green algae, some authors are starting to include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid (spherical), and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae, many of which live most of their lives as single cells, while other species form coenobia (colonies), long filaments ...
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Photobiont
A lichen ( , ) is a hybrid colony of algae or cyanobacteria living symbiotically among filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology. .
Lichens are the lifeform that first brought the term symbiosis (as ''Symbiotismus'') into biological context. Lichens have since been recognized as important actors in and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lich ...
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Epiphytic Fungus
An epiphytic fungus is a fungus that grows upon, or attached to, a living plant. The term epiphytic derives from the Greek ''epi-'' (meaning 'upon') and ''phyton'' (meaning 'plant'). Examples Many examples of epiphytic microorganisms exist. The ergoline alkaloids found in ''Convolvulaceae'' are produced by a seed-transmitted epiphytic clavicipitaceous fungus . See also * Foliicolous, lichens or bryophytes that grow on leaves * Epiphyte * Endosymbiont * Epilith, an organism that grows in a rock * Epibiont, an organism that grows on another life form * Epiphytic bacteria * Mycorrhiza A mycorrhiza (; , mycorrhiza, or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, the plant root system and its surroundings. Mycorrhizae play ... References Mycology {{mycology-stub ...
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Ascocarps
An ascocarp, or ascoma (: ascomata), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of ...
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Crustose Lichen
Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cortex (botany), cortex layer, an algal layer, and a medulla. The upper cortex layer is differentiated and is usually pigmented. The algal layer lies beneath the cortex. The medulla fastens the lichen to the substrate and is made up of Fungus, fungal hyphae. The surface of crustose lichens is characterized by branching cracks that periodically close in response to climatic variations such as alternate wetting and drying regimes. Subtypes * Powdery – considered as the simplest subtype due to the absence of an organized thallus. :The thallus appears powdery. :E.g. Genera ''Lepraria'', ''Vezdaea'' * Endolithic – grows inside the rock, usually in interstitial spaces between mineral grains. The :upper cortex is usually d ...
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Thallus
Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thalloidal, thalliform, thalline, or thallose. Even though thalli do not have organized and distinct parts ( leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where th ...
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Lichens
A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology. .
Lichens are the lifeform that first brought the term symbiosis (as ''Symbiotismus'') into biological context. Lichens have since been recognized as important actors in nutrient cycling and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lichens have properties different from those of their component organisms. They come in man ...
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Monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A '' polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping ...
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Ribosomal DNA
The ribosomal DNA (rDNA) consists of a group of ribosomal RNA encoding genes and related regulatory elements, and is widespread in similar configuration in all domains of life. The ribosomal DNA encodes the non-coding ribosomal RNA, integral structural elements in the assembly of ribosomes, its importance making it the most abundant section of RNA found in cells of eukaryotes. Additionally, these segments include regulatory sections, such as a promoter specific to the RNA polymerase I, as well as both transcribed and non-transcribed spacer segments. Due to their high importance in the assembly of ribosomes for protein biosynthesis, the rDNA genes are generally highly conserved in molecular evolution. The number of copies can vary considerably per species. Ribosomal DNA is widely used for phylogenetic studies. Structure The ribosomal DNA includes all genes coding for the non-coding structural ribosomal RNA molecules. Across all domains of life, these are the structural ...
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Mitochondrial DNA
Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in a eukaryotic cell; most of the DNA is in the cell nucleus, and, in plants and algae, the DNA also is found in plastids, such as chloroplasts. Mitochondrial DNA is responsible for coding of 13 essential subunits of the complex oxidative phosphorylation (OXPHOS) system which has a role in cellular energy conversion. Human mitochondrial DNA was the first significant part of the human genome to be sequenced. This sequencing revealed that human mtDNA has 16,569 base pairs and encodes 13 proteins. As in other vertebrates, the human mitochondrial genetic code differs slightly from nuclear DNA. Since animal mtDNA evolves faster than nuclear genetic markers, it represents a mainstay of phylogenetics and evolutionary biology. It als ...
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