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Enaliosuchus Macrospondylus
''Enaliosuchus'' is a dubious genus of extinct marine crocodyliform within the family Metriorhynchidae that lived during the Valanginian stage of the Early Cretaceous. It is known from fossil remains found in France and Germany and it was first described in 1883. Only the type species is known, which is ''E. macrospondylus''. Etymology The name ''Enaliosuchus'' means "Marine crocodile", and is derived from the Greek ''Enalios''- ("marine") and -''suchos'' ("crocodile"). Discovery and naming The holotype, specimen MB.R.1943.1–16, consists of an atlas-axis complex, three post-axial cervical vertebrae, several dorsal vertebrae, a caudal vertebra, an incomplete femur and a fragmentary sacral rib, and it was discovered in the Stadthagen Formation in Germany. The species ''Enaliosuchus macrospondylus'' was then named for this specimen by Koken (1883). Isolated teeth were subsequently found across Germany and have been assigned to ''Enaliosuchus'', and a partial specimen ( ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian–Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Onto ...
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France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlantic, North Atlantic, the French West Indies, and List of islands of France, many islands in Oceania and the Indian Ocean, giving it Exclusive economic zone of France, one of the largest discontiguous exclusive economic zones in the world. Metropolitan France shares borders with Belgium and Luxembourg to the north; Germany to the northeast; Switzerland to the east; Italy and Monaco to the southeast; Andorra and Spain to the south; and a maritime border with the United Kingdom to the northwest. Its metropolitan area extends from the Rhine to the Atlantic Ocean and from the Mediterranean Sea to the English Channel and the North Sea. Its Regions of France, eighteen integral regions—five of which are overseas—span a combined area of and hav ...
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Monophyly
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A ''polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping are ...
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Phylogenetic
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history. The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question. In addition to their use for inferring phylogenetic pa ...
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Ichthyosaurs
Ichthyosauria is an taxonomy (biology), order of large extinction, extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides. Ichthyosaurians thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago (Myr, Ma) and at least one species survived until about 90 million years ago, into the Late Cretaceous. During the Early Triassic Geologic time scale, epoch, ichthyosaurs and other Ichthyosauromorpha, ichthyosauromorphs evolved from a group of unidentified land reptiles that returned to the sea, in a development similar to how the mammalian land-dwelling ancestors of modern-day dolphins and whales returned to the sea millions of years later, which they gradually came to resemble in a case of convergent evolution. Ichthyosaurians were particularly abundant in the Late Triassic and Early Jurassic periods, until they were replaced as the top ...
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Plesiosaur
The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian stage, about 203 million years ago. They became especially common during the Jurassic Period, thriving until their disappearance due to the Cretaceous–Paleogene extinction event at the end of the Cretaceous Period, about 66 million years ago. They had a worldwide oceanic distribution, and some species at least partly inhabited freshwater environments. Plesiosaurs were among the first fossil reptiles discovered. In the beginning of the nineteenth century, scientists realised how distinctive their build was and they were named as a separate order in 1835. The first plesiosaurian genus, the eponymous ''Plesiosaurus'', was named in 1821. Since then, more than a hundred valid species have been described. In the early twenty-first cent ...
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Mesozoic
The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian reptiles such as the dinosaurs, and of Gymnosperm, gymnosperms such as cycads, ginkgoaceae and Araucariaceae, araucarian conifers; a hot Greenhouse and icehouse earth, greenhouse climate; and the tectonic break-up of Pangaea. The Mesozoic is the middle of the three eras since Cambrian explosion, complex life evolved: the Paleozoic, the Mesozoic, and the Cenozoic. The era began in the wake of the Permian–Triassic extinction event, the largest mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs, Pterosaur, pterosaurs, Mosasaur, mosasaurs, and Plesiosaur, plesiosaurs. The Mesozoic was a time of significant tectonic, climatic, an ...
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Carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ... whose nutrition and energy requirements are met by eating, consumption of animal tissue (biology), tissues (mainly muscle, adipose tissue, fat and other soft tissues) as food, whether through predation or scavenger, scavenging. Nomenclature Mammal order The technical term for mammals in the order (biology), order Carnivora is ''carnivoran'', and they are so-named because most member species in the group have a carnivorous diet, but the similarity of the name of the order and the name of the diet causes confusion. Many but not all carnivorans are meat eaters; a few, such as the fe ...
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Ernst Koken
Ernst Friedrich Rudolph Karl Koken and from 1907 as von Koken (29 May 1860 – 21 November 1912) was a German geologist and paleontologist. He served as a professor of paleontology at the University of Tübingen. Life and work Koken was born in Braunschweig where his father Hermann was a clerk. After local studies he went to the University of Göttingen with studies in Zurich and Berlin. He received a doctorate in 1884 with studies under Wilhelm Dames and Ernst Beyrich. He completed his habilitation as an assistant to Beyrich and became a lecturer at the University of Berlin in 1888. In 1891 he joined the University of Königsberg as a professor to succeed Wilhelm Branco and in 1895 he moved to the University of Tübingen to succeed Branco again, and where he worked until his death. He was involved in establishing the geological and paletontological institute and establishing its collections. His students included Friedrich von Huene. Koken studied Cretaceous dinosaurs from Ger ...
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Stadthagen Formation
The Stadthagen Formation is a geological formation located in Hanover, Germany, dating to the Early Cretaceous (Valanginian). The formation contains fossils of reptiles and fish, suggesting that the formation was once a shallow marine setting.''Pêgas, R.V., Holgado, B., Leal, M.E.C., (2019). "''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids", ''Historical Biology'', 1–15. doi:10.1080/08912963.2019.1690482'' History The Stadthagen Formation outcrops in the Engelbostel and Sachsenhagen clay pits, which operated as brickworks A brickworks, also known as a brick factory, is a factory for the manufacturing of bricks, from clay or shale. Usually a brickworks is located on a clay bedrock (the most common material from which bricks are made), often with a clay pit, quar ... from 1904 to 1986. After the brickworks closed in 1986, most of the quarry ar ...
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Enaliosuchus Vertebra Holotype
''Enaliosuchus'' is a dubious genus of extinct marine crocodyliform within the family Metriorhynchidae that lived during the Valanginian stage of the Early Cretaceous. It is known from fossil remains found in France and Germany and it was first described in 1883. Only the type species is known, which is ''E. macrospondylus''. Etymology The name ''Enaliosuchus'' means "Marine crocodile", and is derived from the Greek ''Enalios''- ("marine") and -''suchos'' ("crocodile"). Discovery and naming The holotype, specimen MB.R.1943.1–16, consists of an atlas-axis complex, three post-axial cervical vertebrae, several dorsal vertebrae, a caudal vertebra, an incomplete femur and a fragmentary sacral rib, and it was discovered in the Stadthagen Formation in Germany. The species ''Enaliosuchus macrospondylus'' was then named for this specimen by Koken (1883). Isolated teeth were subsequently found across Germany and have been assigned to ''Enaliosuchus'', and a partial specimen ( ...
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