Zosterophylls
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Zosterophylls
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxyl ...
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Sawdoniales
The Sawdoniales are an Order (biology), order or plesion of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the Lycopodiopsida, living lycophytes. The group has been divided up in various ways. In their major Cladistics, cladistic study of early land plants, Kenrick and Crane placed most of the zosterophylls in the Sawdoniales (which they treated as a plesion). Like other zosterophylls, members of the Sawdoniales bore lateral, reniform sporangia. They branched dichotomously, and grew at the ends by unrolling (vernation, circinate vernation). Some had smooth stems, others were covered in small spines; fungal bodies have been reported in some spines. Taxonomy In 1997, Kenrick and Crane placed most of the zosterophylls in the plesion Sawdoniales, characterizing the group as having "marked bilateral symmetry". Their summary cladogram did not resolve the taxa within the Sawdoniales, other than placing ''Zos ...
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Hicklingia
''Hicklingia'' is a genus of extinct plants of the Middle Devonian (around ). Compressed specimens were first described in 1923 from the Old Red Sandstone of Scotland., cited in Initially the genus was placed in the "rhyniophytes", but this group is defined as having terminal sporangia (spore-forming organs), and later work showed that the sporangia of ''Hicklingia'' were lateral rather than strictly terminal, so that it is now regarded as having affinities with the zosterophylls. Description The sporophyte had a tufted growth habit, with narrow leafless stems (axes) up to 17 cm high which branched dichotomously. Sporangia were borne on short stalks (up to 3 mm), on all sides of the stem and also terminally. There are oval scars on specimens where the stalks are presumed to have broken off. The lateral sporangia were closely adpressed to the stem. The effect is of a 'spike' of sporangia which terminates some stems. The sporangia opened via slits, but these did not ha ...
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Zosterophyllum
''Zosterophyllum'' was a genus of Silurian-Devonian vascular plant, vascular land plants with naked branching axes on which usually kidney-shaped sporangia were arranged in lateral positions. It is the type genus for the group known as zosterophylls, thought to be part of the lineage from which Lycopodiopsida, modern lycophytes evolved. More than 20 species have been described. Description The diagnostic features of the genus have changed since its first description in 1892, as the original species (''Zosterophyllum myretonianum'') has become better known, and as other species have been discovered. ''Zosterophyllum'' is a vascular plant. The axes (stems) are naked, lacking leaves or outgrowths ("enations"). When branching occurs, the branches are either isotomous (equally sized) or pseudomonopodial (one branch is larger than the other but still clearly involves division of the original axis rather a distinct side growth). The Sporangium, sporangia are upright on short stalks. In ...
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Lycophyte
The lycophytes, when broadly circumscribed, are a group of vascular plants that include the clubmosses. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineages of extant (living) vascular plants; the group contains extinct plants that have been dated from the Silurian (ca. 425 million years ago). Lycophytes were some of the dominating plant species of the Carboniferous period, and included the tree-like Lepidodendrales, some of which grew over in height, although extant lycophytes are relatively small plants. The scientific names and the informal English names used for this group of plants are ambiguous. For example, "Lycopodiophyta" and the shorter "Lycophyta" as well as the informal "lycophyte" may be used to include the extinct zosterophylls or to exclude them. Description Lycophytes reproduce by spores and have alternation of generations in which (like other vascular plants) the sporop ...
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Sawdonia Ornata
Sawdonia is an extinct genus of early vascular plants, known from the Upper Silurian to the Lower Carboniferous (). ''Sawdonia'' is best recognized by the large number of spikes ( enations) covering the plant. These are vascular plants that do not have vascular systems in their enations. The first species of this genus (''Sawdonia ornata'') was described in 1859 by Sir J. William Dawson and, was originally attributed to the genus ''Psilophyton''. He named this plant ''Psilophyton princeps''. In 1971 Francis Hueber proposed a new genus for this species due to its "Divergent technical characters from the generic description for ''Psilophyton''." The holotype used for description is Dawson Collection Number 48, pro parte, Museum Specimen Number 3243. (See Dawson 1871, Plate IX, fig 101.) Sir J. William Dawson Collection, Peter Redpath Museum, McGill University, Montreal, Quebec, Canada. Morphology These plants are described by Hueber as having monopodially branched stems, that ar ...
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Xylem
Xylem is one of the two types of transport tissue (biology), tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports plant nutrition, nutrients. The word ''xylem'' is derived from the Ancient Greek word (), meaning "wood"; the best-known wood organism is plants, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cell (biology), cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Wood also contains two other type of cells: Ground tissue#Parenchyma, parenchyma and ground tissue#Fibres, fibers. Xylem can be found: * in vascular bundles, present in ...
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Dichotomous Branching
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary of leaf morphology. For other related terms, see Glossary of phytopathology, Glossary of lichen terms, and List of Latin and Greek words commonly used in systematic names. A B ...
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Vernation
Vernation or leafing is the formation of new leaves or fronds. In plant anatomy, it is the arrangement of leaves in a bud. In pine species, new leaves are short and encased in sheaths. Each leaf bundle consists of two to five needles. All the leaves on one section of branch grow in length together. In cabbage species, new leaves are folded over, each covered by the previous leaf. Name The term ''vernation'' is borrowed from New Latin , the act of being verdant or flourishing (). It is cognate with (Latin for " spring") and ("vernal"). Circinate vernation Circinate vernation is the manner in which most fern fronds emerge. As the fern frond is formed, it is tightly curled so that the tender growing tip of the frond (and each subdivision of the frond) is protected within a coil. At this stage it is called a crozier (after the shepherd's crook) or fiddlehead (after the scrollwork at the top of a violin). As the lower parts of the frond expand and toughen up, they begin ...
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Stele (biology)
In a vascular plant, the stele is the central part of the root or Plant stem, stem containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue (pith) and a pericycle, which, if present, defines the outermost boundary of the stele. Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by France, French botanists Phillippe Édouard Léon van Tieghem, P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the Shoot (botany), shoot and root, and for discussing the evolution of vascular plant morphology. Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele. Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs. Proto ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta. Angiosperms are by far the most diverse group of Embryophyte, land plants with 64 Order (biology), orders, 416 Family (biology), families, approximately 13,000 known Genus, genera and 300,000 known species. They include all forbs (flowering plants without a woody Plant stem, stem), grasses and grass-like plants, a vast majority of broad-leaved trees, shrubs and vines, and most aquatic plants. Angiosperms are distinguished from the other major seed plant clade, the gymnosperms, by having flowers, xylem consisting of vessel elements instead of tracheids, endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the commo ...
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Xylem Development
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from the Ancient Greek word (), meaning "wood"; the best-known wood organism is plants, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Wood also contains two other type of cells: parenchyma and fibers. Xylem can be found: * in vascular bundles, present in non-woody plants and non-woody parts of woody plants * in secondary xylem, laid down by a merist ...
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