Opuntioideae
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Opuntioideae
Opuntioideae is a subfamily of the cactus family, Cactaceae. It contains 15 genera divided into five tribes. The subfamily encompasses roughly 220–250 species, and is geographically distributed throughout the New World from Canada, to Argentina. Members of this subfamily have diverse habits, including small ..., including small geophytes, hemispherical cushions, shrub">geophytes">..., including small geophytes, hemispherical cushions, shrubs, trees, and columnar cacti consisting of indeterminate branches or determinate terete or spherical segments. Description Synapomorphies of Opuntioideae include small deciduous, barbed spines called glochids born on areoles and a bony aril surrounding a campylotropous ovule (inverted and curved, such that the micropyle almost meets the funiculus). Other prominent morphological characters for this subfamily are presence of cylindrical, caducous leaves that tend to be shed by maturity and the sectioning of the stem into joints or pads known ...
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Cactus
A cactus (: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word ''cactus'' derives, through Latin, from the Ancient Greek word (''káktos''), a name originally used by Theophrastus for a spiny plant whose identity is now not certain. Cacti occur in a wide range of shapes and sizes. They are native to the Americas, ranging from Patagonia in the south to parts of western Canada in the north, with the exception of ''Rhipsalis baccifera'', which is also found in Africa and Sri Lanka. Cacti are adapted to live in very dry environments, including the Atacama Desert, one of the driest places on Earth. Because of this, cacti show many adaptations to conserve water. For example, almost all cacti are succulents, meaning they have thickened, fleshy parts adapted to store water. Unlike many other succulents, the stem is the only part of most cacti ...
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Areole
In botany, areoles are small light- to dark-colored bumps on cactus, cacti out of which grow clusters of Thorns, spines, and prickles, spines. Areoles are important diagnostic features of cactus, cacti, and identify them as a family distinct from other succulent plants. Gordon Douglas Rowley, Gordon Rowley - What is an Areole The spines are not easily detachable, but on certain cacti, members of the subfamily Opuntioideae, smaller, detachable bristles, glochids, also grow out of the areoles and afford additional protection. Areoles represent highly specialized branches on cacti. They evolved as abortive branch buds while their spines evolved as vestigial leaves. In branched cacti, such as Opuntioidiae and the saguaro, new branches grow from areoles, because that is where the buds are. The development of the areole seems to have been an important element in the adaptation of cacti to niches in desert ecology. Some of the Opuntioideae have spines, as well as glochids, on their a ...
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Glochid
Glochids or glochidia (: "glochidium") are hair-like spines or short prickles, generally barbed, found on the areoles of Cactus, cacti in the sub-family ''Opuntioideae''. Cactus glochids easily detach from the plant and lodge in the skin, causing irritation upon contact. The tufts of glochids in the areoles nearly cover the stem surfaces of some cactus species, each tuft containing hundreds of glochids; this may be in addition to, or instead of, the larger, more conspicuous cactus Spine (botany), spines, which do not readily detach and are not generally barbed. Mechanical irritant dermatitis Reaction to the glochidia Most cacti possess spines, some large enough to cause serious wounds. Glochids however, though smaller, commonly induce more troublesome, more persistent, dermatological manifestations in humans. Though minute, glochids commonly are barbed and once they have penetrated the skin barbed glochids are practically impossible to dislodge without leaving scraps of fore ...
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Cylindropuntia Ramosissima
''DCylindropuntia ramosissima'' is a species of cactus known by the common names Diamond cholla and Branched pencil cholla. Distribution ''Cylindropuntia ramosissima'' is native to the Mojave and Sonoran Deserts of the Southwestern United States, California, and Northwestern Mexico, and to Baja California and its Islas San Benito. Description ''Cylindropuntia ramosissima'' is a decumbent or erect and treelike cactus which can approach 2 meters or 6 feet in maximum height. It has many narrow branches made up of cylindrical segments, green in color drying gray, the surface divided into squarish, flat tubercles with few or no spines, or often with a single long, straight spine. The flower is small and orange, pink or brownish in color. The fruit is a small, dry, spiny body up to 2 centimeters long. ''Cylindropuntia ramosissima'' is a host to several species of moths, most notably the Sphinx Moth The Sphingidae are a family of moths commonly called sphinx moths, also colloqui ...
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Tubercle
In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection, but it has slightly different meaning depending on which family of plants or animals it is used to refer to. In the case of certain orchids and cacti, it denotes a round nodule, small eminence, or warty outgrowth found on the lip. They are also known as podaria (singular ''podarium''). When referring to some members of the pea family, it is used to refer to the wart-like excrescences that are found on the roots. In fungi In mycology, a tubercle is used to refer to a mass of hyphae from which a mushroom is made. In animals When it is used in relation to certain dorid nudibranchs such as '' Peltodoris nobilis'', it means the nodules on the dorsum of the animal. The tubercles in nudibranchs can present themselves in different way ...
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Epidermis (botany)
The epidermis (from the Greek language, Greek ''ἐπιδερμίς'', meaning "over-skin") is a single layer of cells that covers the leaf, leaves, flowers, roots and Plant stem, stems of plants. It forms a boundary between the plant and the external environment. The epidermis serves several functions: it protects against water loss, regulates gas exchange, secretes metabolic compounds, and (especially in roots) absorbs water and mineral nutrients. The epidermis of most leaves shows Anatomical terms of location#Dorsal and ventral, dorsoventral anatomy: the upper (adaxial) and lower (abaxial) surfaces have somewhat different construction and may serve different functions. Woody stems and some other stem structures such as potato tubers produce a secondary covering called the periderm that replaces the epidermis as the protective covering. Description The epidermis is the outermost cell layer of the primary plant body. In some older works the cells of the leaf epidermis have been ...
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Trichome
Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant is an indumentum, and the surface bearing them is said to be Leaf#Surface, pubescent. Algal trichomes Certain, usually filamentous, algae have the terminal cell (biology), cell produced into an elongate hair-like structure called a trichome. The same term is applied to such structures in some cyanobacteria, such as ''Spirulina (dietary supplement), Spirulina'' and ''Oscillatoria''. The trichomes of cyanobacteria may be unsheathed, as in ''Oscillatoria'', or sheathed, as in ''Calothrix''. These structures play an important role in preventing soil erosion, particularly in cold desert climates. The filamentous sheaths form a persistent sticky network that helps maintain soil structure. Plant trichomes Plant trichomes have many diff ...
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Mucilage
Mucilage is a thick gluey substance produced by nearly all plants and some microorganisms. These microorganisms include protists which use it for their locomotion, with the direction of their movement always opposite to that of the secretion of mucilage. It is a polar glycoprotein and an exopolysaccharide. Mucilage in plants plays a role in the storage of water and food, seed germination, and thickening membranes. Cacti (and other succulents) and flax seeds are especially rich sources of mucilage. Occurrence Exopolysaccharides are the most stabilising factor for microaggregates and are widely distributed in soils. Therefore, exopolysaccharide-producing "soil algae" play a vital role in the ecology of the world's soils. The substance covers the outside of, for example, unicellular or filamentous green algae and cyanobacteria. Amongst the green algae especially, the group Volvocales are known to produce exopolysaccharides at a certain point in their life cycle. It occu ...
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Internodes
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, engages in photosynthesis, stores nutrients, and produces new living tissue. The stem can also be called the culm, halm, haulm, stalk, or thyrsus. The stem is normally divided into nodes and internodes: * The nodes are the points of attachment for leaves and can hold one or more leaves. There are sometimes axillary buds between the stem and leaf which can grow into branches (with leaves, conifer cones, or flowers). Adventitious roots (e.g. brace roots) may also be produced from the nodes. Vines may produce tendrils from nodes. * The internodes distance one node from another. The term " shoots" is often confused with "stems"; "shoots" generally refers to new fresh plant growth, including both stems and other structures like leaves or flowers. In most ...
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Phloem
Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is called translocation. In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word (''phloiós''), meaning "bark". The term was introduced by Carl Nägeli in 1858. Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem. Protophloem eventually becomes obliterated once it connects to the durable phloem in mature organs, the metaphloem. Further, secondary phloem is formed during the thickening of stem structures. Structure Phloem tissue consists of conducting cells, generally called sieve elements, parenchyma cells, including both specialized companion cells or albuminous cells and unspecialized cells and supportive cells, such ...
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Cladode
Phylloclades and cladodes are flattened, photosynthesis, photosynthetic shoots, which are usually considered to be modified branches. The two terms are used either differently or interchangeably by different authors. ''Phyllocladus'', a genus of conifer, is named after these structures. Phylloclades/cladodes have been identified in fossils dating from as early as the Permian. Definition and morphology The term "phylloclade" is from the Neo-Latin ''phyllocladium'', itself derived from Greek language, Greek ''phyllo'', leaf, and ''klados'', branch. Definitions of the terms "phylloclade" and "cladode" vary. All agree that they are flattened structures that are photosynthetic and resemble leaf-like branches. In one definition, phylloclades are a subset of cladodes, namely those that greatly resemble or perform the function of leaves, as in Butcher's broom (''Ruscus aculeatus'') as well as ''Phyllanthus'' and some ''Asparagus'' species. By an alternative definition, cladodes are di ...
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