Liverworts
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Liverworts
Liverworts are a group of non-vascular plant, non-vascular embryophyte, land plants forming the division Marchantiophyta (). They may also be referred to as hepatics. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information. The division name was derived from the genus name ''Marchantia'', named after his father by French botanist Jean Marchant. It is estimated that there are about 9000 species of liverworts. Some of the more familiar species grow as a flattened leafless thallus, but most species are leafy with a form very much like a flattened moss. Leafy species can be distinguished from the apparently similar mosses on the basis of a number of features, including their single-celled rhizoids. Leafy liverworts also differ from most (but not all) mosses in that their leaves never have a costa (botany), costa (present in many mosses) and may bear marginal cilia (botany), cilia (very rare i ...
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Metzgeriales
Metzgeriales is an order of liverworts. The group is sometimes called the simple thalloid liverworts: " thalloid" because the members lack structures resembling stems or leaves, and "simple" because their tissues are thin and relatively undifferentiated. All species in the order have a small gametophyte stage and a smaller, relatively short-lived, spore-bearing stage. Although these plants are almost entirely restricted to regions with high humidity or readily available moisture, the group as a whole is widely distributed, and occurs on every continent except Antarctica. Description Members of the Metzgeriales typically are small and thin enough to be translucent, with most of the tissues only a single cell layer in thickness. Because these plants are thin and relatively undifferentiated, with little evidence of distinct tissues, the Metzgeriales are sometimes called the "simple thalloid liverworts". There is considerable diversity in vegetative structure of the Metzger ...
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Marchantiales
Marchantiales is an order of thallose liverworts (also known as "complex thalloid liverworts") that includes species like '' Marchantia polymorpha'', a widespread plant often found beside rivers, and '' Lunularia cruciata'', a common and often troublesome weed in moist, temperate gardens and greenhouses. As in other bryophytes, the gametophyte generation is dominant, with the sporophyte existing as a short-lived part of the life cycle, dependent upon the gametophyte. The genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ... '' Marchantia'' is often used to typify the order, although there are also many species of '' Asterella'' and species of the genus '' Riccia'' are more numerous. The majority of genera are characterized by the presence of (a) special stalked vertical bran ...
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Mosses
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically tall, though some species are much larger. ''Dawsonia'', the tallest moss in the world, can grow to in height. There ar ...
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Haplomitriopsida
Haplomitriopsida is a Class (biology), class of Marchantiophyta, liverworts comprising fifteen species in three genus, genera. Recent cladistics, cladistic analyses of cell nucleus, nuclear, mitochondrial, and plastid gene sequences place this monophyly, monophyletic group as the basal sister group to all other liverworts. The group thus provides a unique insight into the early evolution of liverworts in particular and of embryophyte, land plants in general. Description Plants of ''Treubia'' grow as a prostrate leafy thallus (tissue), thallus. The wikt:bifid, bifid leaves extend like wings on either side of the midrib, or may be folded upwards and pressed close together, giving the plants a ruffled appearance. By contrast, ''Haplomitrium'' grows as a wikt:subterranean, subterranean rhizome with erect leafy stems. The thin, rounded leaves are arranged around the upright stems, giving the appearance of a soft moss. The species ''Haplomitrium ovalifolium'' of Australia often has b ...
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Lunulariales
''Lunularia'' is a genus of liverworts whose only species is ''Lunularia cruciata'', the crescent-cup liverwort. ''Lunularia'' is either the only genus in the order Lunulariales, or may be placed in the order Marchantiales. The name, from Latin ''luna'', moon, refers to the moon-shaped gemma cups. Description ''Lunularia cruciata'' grows large, dichotomously branched green thalli with crescent shaped gemma cups containing disc-like gemmae. This is a unique morphological characteristic not possessed by other thalloid liverworts. Its thallus surface is shiny, faintly lined, and is dotted with tiny air pores. When dried the thallus turns yellowish in color and its margin rolls inward. As in other liverworts, the main plant body or thallus is a haploid gametophyte. The antheridia of ''L. cruciata'' develops in early spring, the archegonia develops in spring and sporophytes develop in late summer. However, records of sporophyte developments and sexual reproduction are rare a ...
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Marchantiopsida
Marchantiopsida is a class of liverworts within the phylum Marchantiophyta. The species in this class are known as complex thalloid liverworts. The species in this class are widely distributed and can be found worldwide. Complex oil bodies are only found in the gametophyte. Taxonomy * Blasiidae He-Nygrén et al. 2006 ** Blasiales Stotl. & Crand.-Stotl. 2000 *** Blasiaceae H.Klinggr. 1858 *** †Treubiitaceae Schuster 1980 * Marchantiidae Engl. 1893 sensu He-Nygrén et al. 2006 ** Lunulariales H.Klinggr. 2006 *** Lunulariaceae H.Klinggr. 1858 ** Marchantiales Limpr. 1877 (complex thalloids) *** Aytoniaceae Cavers 1911 [Rebouliaceae; Grimaldiaceae] *** Cleveaceae Cavers 1911 [Sauteriaceae] *** Conocephalaceae Müll.Frib. ex Grolle 1972 *** Corsiniaceae Engl. 1892 *** Cyathodiaceae Stotler & Crand.-Stotl. 2000 *** Dumortieraceae Long 2006 *** Exormothecaceae Müll.Frib. ex Grolle 1972 *** Marchantiaceae Lindl. 1836 *** Monocleaceae A.B.Frank 1877 *** Monosoleniaceae Inoue 1966 ...
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Hornworts
Hornworts are a group of Non-vascular plant, non-vascular Embryophytes (land plants) constituting the division Anthocerotophyta (). The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information; the flattened, green plant body of a hornwort is the gametophyte stage of the plant. Hornworts may be found worldwide, though they tend to grow only in places that are damp or humid. Some species grow in large numbers as tiny weeds in the soil of gardens and cultivated fields. Large tropical and sub-tropical species of ''Dendroceros'' may be found growing on the bark of trees. The total number of species is still uncertain. While there are more than 300 published species names, the actual number could be as low as 100–150 species. Description Like all bryophytes, the dominant life phase of a hornwort is the h ...
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Blasiales
Blasiales is an order of liverworts with a single living family and two species. The order has traditionally been classified among the Metzgeriales, but molecular cladistics suggests a placement at the base of the Marchantiopsida. Taxonomy * Blasiales Stotler & Crandall-Stotler 2000 ** Blasiaceae von Klinggräff 1858 *** ''Blasia'' Linnaeus 1753 **** ''Blasia pusilla'' Linnaeus 1753 *** ''Cavicularia'' Stephani 1897 non Pavesi 1881 **** ''Cavicularia densa'' Stephani 1897 ** †Treubiitaceae Schuster 1980 *** †''Treubiites ''Treubiites kidstonii'' is a fossil species of liverworts in the family Treubiitaceae. The only known fossils come from Late Carboniferous deposits of Shropshire, England England is a Countries of the United Kingdom, country that is pa ...'' Schuster 1966 **** †''Treubiites kidstonii'' (Walton 1925) Schuster 1966 References External links Liverwort Tree of Life Liverwort orders {{Bryophyte-stub ...
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Jungermanniales
Jungermanniales is the largest Order (biology), order of Marchantiophyta, liverworts. They are distinctive among the liverworts for having thin leaf-like flaps on either side of the stem. Most other liverworts are thalloid, with no leaves. Due to their dorsiventral organization and scale-like, overlapping leaves, the Jungermanniales are sometimes called "scale-mosses". Families of Jungermanniales An updated classification by Söderström et al. 2016 * Cephaloziineae Schljakov [Jamesoniellineae] ** Adelanthaceae Grolle 1972 [Jamesoniellaceae He-Nygrén et al. 2006] ** Anastrophyllaceae Söderström et al. 2010b ** Cephaloziaceae Walter Migula, Migula 1904 ** Cephaloziellaceae Douin 1920 [Phycolepidoziaceae Schuster 1967] ** Lophoziaceae Cavers 1910 ** Scapaniaceae Migula 1904 [Diplophyllaceae Potemk. 1999; Chaetophyllopsaceae Schuster 1960] * Jungermanniineae Schuster ex Stotler & Crandall-Stotler 2000 [Geocalycineae Schuster 1972] ** Acrobolbaceae Hodgson 1962 ** Antheliaceae Sch ...
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Pleuroziales
''Pleurozia'' is the only genus of liverworts in the family Pleuroziaceae, which is now classified in its own order Pleuroziales, but was previously included in a broader circumscription of the Jungermanniales. The genus includes twelve species, and as a whole is both physically distinctive and widely distributed. The lower leaf lobes of ''Pleurozia'' species are fused, forming a closed water sac covered by a movable lid similar in structure to those of the angiosperm genus ''Utricularia''. These sacs were assumed to play a role in water storage, but a 2005 study on '' Pleurozia purpurea'' found that the sacs attract and trap ciliates, much in the same way as ''Utricularia''. Observations of plants ''in situ'' also revealed a large number of trapped prey within the sacs, suggesting that the species in this genus obtain some benefit from a carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour" ...
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Jungermanniopsida
Jungermanniopsida is the largest of three classes within the division Marchantiophyta (liverworts). Phylogeny Based on the work by Novíkov & Barabaš-Krasni 2015. Taxonomy * Jungermanniidae Engler 1893 ** Jungermanniales von Klinggräff 1858 *** Cephaloziineae Schljakov **** Adelanthaceae Grolle 1972 amesoniellaceae He-Nygrén et al. 2006**** Anastrophyllaceae Söderström et al. 2010b **** Cephaloziaceae Migula 1904 **** Cephaloziellaceae Douin 1920 hycolepidoziaceae Schuster 1967**** Lophoziaceae Cavers 1910 **** Scapaniaceae Migula 1904 iplophyllaceae Potemk. 1999; Chaetophyllopsaceae Schuster 1960*** Jungermanniineae Schuster ex Stotler & Crandall-Stotler 2000 **** Acrobolbaceae Hodgson 1962 **** Antheliaceae Schuster 1963 **** Arnelliaceae Nakai 1943 **** Balantiopsidaceae Buch 1955 sotachidaceae**** Blepharidophyllaceae Schuster 2002 **** Calypogeiaceae Arnell 1928 izutaniaceae Furuki & Iwatsuki 1989**** Endogemmataceae Konstantinova, Vilnet & Troits ...
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Calobryales
Calobryales (formerly Haplomitriales) is an order of plants known as liverworts. This order contains one family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ..., Haplomitriaceae, with a single extant genus '' Haplomitrium''. Taxonomy * Order Calobryales Campbell ex Hamlin 1972 aplomitriales Buch ex Schljakov 1972** Family Haplomitriaceae Dědeček 1884 *** Genus †'' Gessella'' Poulsen 1974 *** Genus '' Haplomitrium'' Nees 1833 nom. cons. References External links Information on family Haplomitriaceae Liverwort families {{Bryophyte-stub ...
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