Lithornithidae
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Lithornithidae
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site. Lithornithids had long, slender, bills for probing. They closely resembled modern tinamous, aside from more developed wings. They possessed a rhynchokinetic skull with relatively unfused cranial bones, a weakly fused pygostyle and a splenial. The unguals were more curved than in tinamous and probably allowed better perching in trees. The order Lithornithiformes was erected by Dr. Peter Houde in 1988. Initially, only three genera ('' Lithornis'', '' Paracathartes'', and '' Pseudocrypturus'') and eight named species were included. '' Promusophaga'' (Harrison & Walker, 1977) originally considered ...
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Lithornis
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. ''Lithornis'' is from ancient Greek for 'stone bird', as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are '' Paracathartes'' and '' Pseudocrypturus''. Some researchers consider '' Calciavis'' as a species of ''Lithornis'' (''L. grandei''). Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird '' Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is ...
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Calciavis Grandei
''Calciavis'' is an extinct genus of bird from the Eocene of Wyoming. It is a lithornithid, a member of a lineage of flying palaeognaths, distantly related to modern ratites. Like many other fossils from the Green River Formation, its are exceptionally well preserved, bearing impressions of skin and feathers. Some researchers consider this taxon as a species of ''Lithornis'' (''L. grandei''). Discovery and description The original specimen of ''Calciavis'', AMNH 30578, was found in the Fossil Butte Member of the Green River Formation, Wyoming, dating to around 51 million years ago. It is an exceptionally well preserved specimen, preserving most of the post-cranial skeleton aside from the femur and associated pelvic region, as well as soft-tissue impressions of feathers, skin, foot scales and claw sheaths. Another specimen, AMNH 30560 , was found in the vicinity, composed of a similarly complete most mostly disarticulate skeleton. It is diagnosed by several features of the skul ...
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Lithornis Celetius
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. ''Lithornis'' is from ancient Greek for 'stone bird', as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are '' Paracathartes'' and '' Pseudocrypturus''. Some researchers consider '' Calciavis'' as a species of ''Lithornis'' (''L. grandei''). Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is fr ...
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Pseudocrypturus Cercanaxius
''Pseudocrypturus'' is a genus of extinct paleognathous bird. Three species are known and the type species is ''Pseudocrypturus cercanaxius''. It is a relative of such modern birds as ostriches. It lived in the early Eocene. The holotype fossil is in the collection of the Smithsonian's National Museum of Natural History. It has catalog number USNM 336103. It was collected from the Fossil Butte Member, Green River Formation, Lincoln County, Wyoming.Houde, Peter W. (1988) Taxonomy ''Pseudocrypturus'' means false tinamou. The species name cercanaxius comes from ancient Greek words , tail, and , worthless, in reference to the rudimentary pygostyle Pygostyle is a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main component o ... of this species. Footnotes References * * {{Taxonbar, from=Q7254714 Bird gen ...
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Messel Pit
The Messel Formation is a Formation (geology), geologic formation in Hesse, central Germany, dating back to the Eocene Epoch (geology), epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlies crystalline Variscan basement and its Permian cover (Rotliegend) as well as Eocene volcanic breccias derived from the basement rocks. The formation mainly comprises lacustrine laminated bituminous shale (‘oil shale’) renowned for its content of fossils in exceptional preservation, particularly plants, arthropods and vertebrates (e.g. ''Darwinius masillae''). Messel pit The Messel pit () is a disused quarry near the village of Messel (Landkreis Darmstadt-Dieburg, Hesse) about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserved fossils of the Messel Formation dating from the middle of the Eocene, it has significant geological and scientific importance. Over 1400 taxa o ...
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Fissuravis
''Fissuravis'' ("fissure bird") is a genus of extinct bird from the Paleocene of Germany. A lithornithid, it was closely related to modern ratites, but it was a capable flyer.Gerald Mayr, The Birds from the Paleocene Fissure Filling of Walbeck (Germany), Journal of Vertebrate Paleontology 27(2):394–408, June 2007 Ecology Hailing from the Walbeck Paleocene deposits, it is found amidst a rich avian fauna, which also included the gigantic '' Gastornis'' and the enigmatic ratite '' Remiornis''. Like most lithornithids, it was probably a very competent flyer, its coracoid remnants suggesting powerful flight musculature, and it likely engaged in a similar style of soaring flight to ''Lithornis'' and ''Pseudocrypturus ''Pseudocrypturus'' is a genus of extinct paleognathous bird. Three species are known and the type species is ''Pseudocrypturus cercanaxius''. It is a relative of such modern birds as ostriches. It lived in the early Eocene. The holotype fossil ...''. References ...
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