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''Khaan'' (; from Mongol 'lord') is an extinct genus of oviraptorid dinosaur known from remains found in the Djadochta Formation of Mongolia, dating to the Late Cretaceous (Campanian, 75-71 million years ago). Description ''Khaan'' did not differ much from other oviraptorids. At first, its remains were assigned to " Ingenia", but the ''Khaan'' manual structure, lacking the expansion of the upper third metacarpal, was considered to differ sufficiently from that of "Ingenia" for it to be assigned to its own genus. The oviraptorid diet is disputed, with plants and molluscs having been suggested. Like other oviraptorids, ''Khaan'' was probably at least partially a meat eater, feeding on small vertebrates like mammals, lizards and possibly other small dinosaurs. It was also probably feathered. Discovery The type species ''Khaan mckennai'' was named by James M. Clark ''et al.'' in 2001. The genus name is derived from Mongol ''khaan'', "lord" or "ruler". The specific name hon ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet, species epithet, or epitheton) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Etymology Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the ...
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Heyuannia
''Heyuannia'' ("from Heyuan") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch, in what is now China and Mongolia. It was the first oviraptorid found in China; most others were found in neighbouring Mongolia. Two species are known: ''H. huangi'', named by Lü Junchang in 2002 from the Dalangshan Formation; and ''H. yanshini'', originally named as a separate genus ''Ingenia'' from the Barun Goyot Formation by Rinchen Barsbold in 1981, and later renamed to ''Ajancingenia'' in 2013 due to the preoccupation of ''Ingenia''. The latter name was eventually discarded due to various ethical issues surrounding the author. Discovery and naming ''H. huangi'' The type species, ''Heyuannia huangi'', was named and described by Lü Junchang in 2002. The generic name refers to the city of Heyuan. The specific name (zoology), specific name honours Huang Dong, the director of the Heyuan Museum. The holotype, HYMV1-1, was discovered in Guangdong near Huangsha i ...
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Nemegtomaia
''Nemegtomaia'' is a genus of oviraptorid dinosaur from what is now Mongolia that lived in the Late Cretaceous Period (geology), Period, about 70million years ago. The first specimen was found in 1996, and became the basis of the new genus and species ''N. barsboldi'' in 2004. The original genus name was ''Nemegtia'', but this was changed to ''Nemegtomaia'' in 2005, as the former name was preoccupied. The first part of the generic name refers to the Nemegt Basin, where the animal was found, and the second part means "good mother", in reference to the fact that oviraptorids are known to have brooded their eggs. The Specific name (zoology), specific name honours the palaeontologist Rinchen Barsbold. Two more specimens were found in 2007, one of which was found on top of a nest with eggs, but the dinosaur had received its genus name before it was found associated with eggs. ''Nemegtomaia'' is estimated to have been around 2 m (7 ft) in length, and to have weighed 40 k ...
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Machairasaurus
''Machairasaurus'' is a genus of oviraptorid dinosaur which was found in the Bayan Mandahu Formation, China, dating to the late Cretaceous period. Discovery During the Sino-Canadian expeditions of 1988 and 1990 some skeletons of unknown oviraptorosaurians were discovered by Philip J. Currie in Inner Mongolia. Based on two of these a new genus was named and described by Nicholas Longrich, Philip Currie, and Dong Zhiming in 2010 with the type species ''Machairasaurus leptonychus''. The generic name is derived from Greek ('), "short scimitar". The specific name is derived from Greek ('), "slender", and ('), "claw". The species name as a whole refers to the sabre-like claws of the hand. The holotype, IVPP V15979, was found in layers of the Bayan Mandahu dating from the late Campanian. It mainly consists of a left front limb, including the lower end of the lower arm, two carpal bones, and a complete hand.Some fragmentary foot elements are also known. The other find is the parat ...
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Citipati (dinosaur)
''Citipati'' (; meaning "funeral pyre lord") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadochta Formation, where the first remains were collected during the 1990s. The genus and type species ''Citipati osmolskae'' were named and described in 2001. A second species from the adjacent Zamyn Khondt locality may also exist. ''Citipati'' is one of the best-known oviraptorids thanks to a number of well-preserved specimens, including individuals found in brooding positions atop nests of eggs, though most of them were initially referred to the related ''Oviraptor''. These nesting specimens have helped to solidify the link between non-avian dinosaurs and birds. ''Citipati'' was among the largest oviraptorids; it is estimated to have been around in length and to have weighed . Its skull was highly pneumatized, short, and had a characteristic ...
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Rinchenia
''Rinchenia'' (named after Byambyn Rinchen) is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch in what is now Mongolia, Nemegt Formation, around 70 million years ago. The type species, type and only known species, ''Rinchenia mongoliensis'', was originally classified as a species within the genus ''Oviraptor'' (named ''Oviraptor mongoliensis''), but a subsequent reexamination found differences significant enough to warrant a separate genus. The name ''Rinchenia'' was coined for this new genus, though not formally described in detail. History of discovery During 1984, a nearly complete oviraptorid skeleton was discovered at the Altan Uul II (or Altan Ula II) locality of the highly fossiliferous Nemegt Formation, Gobi Desert. This newly collected specimen, MPC-D 100/32-A, included the skull and lower jaws in their entirety, nearly complete vertebral column, forelimbs with shoulder girdle, and partial hindlimbs with pelvic girdle. Later on, Mongol ...
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Oviraptor
''Oviraptor'' (; ) is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period. The first remains were collected from the Djadokhta Formation of Mongolia in 1923 during a paleontological expedition led by Roy Chapman Andrews, and in the following year the genus and type species ''Oviraptor philoceratops'' were named by Henry Fairfield Osborn. The genus name refers to the initial thought of egg-stealing habits, and the specific name was intended to reinforce this view indicating a preference over ceratopsian eggs. Despite the fact that numerous specimens have been referred to the genus, ''Oviraptor'' is only known from a single partial skeleton regarded as the holotype, as well as a nest of about fifteen eggs and several small fragments from a juvenile. ''Oviraptor'' was a rather small feathered oviraptorid, estimated at long with a weight between . It had a wide lower jaw with a skull that likely had a crest. Both upper and lower jaws were toothless ...
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Cladogram
A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other e ...
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Conchoraptor
''Conchoraptor'' (meaning "conch plunderer") is a genus of oviraptorid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt Formation, Nemegt formations of Mongolia. Discovery When first discovered in the Red Beds of Hermiin Tsav of the Early Maastrichtian Nemegt Formation by a Polish-Mongolian expedition in 1971, scientists believed that ''Conchoraptor'' was a juvenile ''Oviraptor'' and that the animal's missing crest would have begun to grow when the animal reached sexual maturity. Further study of multiple skeletons showed that ''Conchoraptor'' belonged to a new genus. The hands of ''Conchoraptor'' were a major reason scientists decided to split it off from ''Oviraptor''. Anatomically the hands seemed to be an evolutionary intermediate between those of ''Ajancingenia'' and ''Oviraptor'', making it obvious that this animal was not a member of a known species. The type species of this new genus, ...
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Khaan Skeletal Headden
''Khaan'' (; from Mongol 'lord') is an extinct genus of oviraptorid dinosaur known from remains found in the Djadochta Formation of Mongolia, dating to the Late Cretaceous (Campanian, 75-71 million years ago). Description ''Khaan'' did not differ much from other oviraptorids. At first, its remains were assigned to " Ingenia", but the ''Khaan'' manual structure, lacking the expansion of the upper third metacarpal, was considered to differ sufficiently from that of "Ingenia" for it to be assigned to its own genus. The oviraptorid diet is disputed, with plants and molluscs having been suggested. Like other oviraptorids, ''Khaan'' was probably at least partially a meat eater, feeding on small vertebrates like mammals, lizards and possibly other small dinosaurs. It was also probably feathered. Discovery The type species ''Khaan mckennai'' was named by James M. Clark ''et al.'' in 2001. The genus name is derived from Mongol ''khaan'', "lord" or "ruler". The specific name hon ...
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