Hominina
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Hominina
The australopithecines (), formally Australopithecina or Hominina, are generally any species in the related genera of ''Australopithecus'' and ''Paranthropus''. It may also include members of ''Kenyanthropus'', ''Ardipithecus'', and '' Praeanthropus''. The term comes from a former classification as members of a distinct subfamily, the Australopithecinae. They are classified within the Australopithecina subtribe of the Hominini tribe. These related species are sometimes collectively termed australopithecines, australopiths, or homininians. They are the extinct, close relatives of modern humans and, together with the extant genus ''Homo'', comprise the human clade. There is no general agreement to whether australopithecines are closest relatives of modern humans, as it has been argued that they are more closely related to extant African apes. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are called Hominina (''see Hominidae; terms "hominids" a ...
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Hominidae
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic Family (biology), family of primates that includes eight Neontology#Extant taxa versus extinct taxa, extant species in four Genus, genera: ''Orangutan, Pongo'' (the Bornean orangutan, Bornean, Sumatran orangutan, Sumatran and Tapanuli orangutan); ''Gorilla'' (the Eastern gorilla, eastern and western gorilla); ''Pan (genus), Pan'' (the chimpanzee and the bonobo); and ''Homo'', of which only Human, modern humans (''Homo sapiens'') remain. Numerous revisions in classifying the great apes have caused the use of the term ''hominid'' to change over time. The original meaning of "hominid" referred only to humans (''Homo'') and their closest extinct relatives. However, by the 1990s humans and other apes were considered to be "hominids". The earlier restrictive meaning has now been largely assumed by the term ''Hominini, hominin'', which comprises all members of the human clade after the split ...
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Hominini
The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the genus ''Gorilla (genus), Gorilla'' (gorillas), which is grouped separately within the subfamily Homininae. The term Hominini was originally introduced by Camille Arambourg (1948), who combined the categories of ''Hominina'' and ''Simiina'' pursuant to John Edward Gray, Gray's classifications (1825). Traditionally, chimpanzees, gorillas and orangutans were grouped together, excluding humans, as pongidae, pongids. Since Gray's classifications, evidence accumulating from genetic phylogeny confirmed that humans, chimpanzees, and gorillas are more closely related to each other than to the orangutan. The orangutans were reassigned to the family Hominidae (great apes), which already included humans; and the gorillas were grouped as a separate tr ...
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Homo
''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively called archaic humans) classified as either ancestral or closely related to modern humans; these include ''Homo erectus'' and ''Homo neanderthalensis''. The oldest member of the genus is ''Homo habilis'', with records of just over 2 million years ago. ''Homo'', together with the genus ''Paranthropus'', is probably most closely related to the species ''Australopithecus africanus'' within ''Australopithecus''.'''' The closest living relatives of ''Homo'' are of the genus ''Pan (genus), Pan'' (chimpanzees and bonobos), with the ancestors of ''Pan'' and ''Homo'' estimated to have diverged around 5.7–11 million years ago during the Late Miocene. ''H. erectus'' appeared about 2 million years ago and spread throughout Africa (deba ...
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Sahelanthropus
''Sahelanthropus'' is an extinct genus of hominid dated to about during the Late Miocene. The type species, ''Sahelanthropus tchadensis'', was first announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in northern Chad. The definitive phylogenetic position of ''Sahelanthropus'' within hominids is uncertain. It was initially described as a possible hominin ancestral to both humans and chimpanzees, but subsequent interpretations suggest that it could be an early member of the tribe Gorillini or a stem-hominid outside the hominins. Examinations on the postcranial skeleton of ''Sahelanthropus'' also indicated that this taxon was not a habitual biped. Taxonomy Discovery Four employees of the Centre National d'Appui à la Recherche (CNAR, National Research Support Center) of the Ministry of Higher Education of the Republic of Chad, three Chadians (Ahounta Djimdoumalbaye, Fanoné Gongdibé and Mahamat Adoum) and one French (Alain Beauvilain) collecte ...
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Tribe (biology)
In biology, a tribe is a taxonomic rank above genus, but below family and subfamily. It is sometimes subdivided into subtribes. By convention, all taxa ranked above species are capitalized, including both tribe and subtribe. In zoology, the standard ending for the name of a zoological tribe is "-ini". Examples include the tribes Caprini (goat-antelopes), Hominini (hominins), Bombini (bumblebees), and Thunnini (tunas). The tribe Hominini is divided into subtribes by some scientists; subtribe Hominina then comprises "humans". The standard ending for the name of a zoological subtribe is "-ina". In botany, the standard ending for the name of a botanical tribe is "-eae". Examples include the tribes Acalypheae and Hyacintheae. The tribe Hyacintheae is divided into subtribes, including the subtribe Massoniinae. The standard ending for the name of a botanical subtribe is "-inae". In bacteriology, the form of tribe names is as in botany, e.g., Pseudomonadeae, based on the ge ...
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Ardipithecus
''Ardipithecus'' is a genus of an extinct hominine that lived during the Late Miocene and Early Pliocene epochs in the Afar Depression, Ethiopia. Originally described as one of the earliest ancestors of humans after they diverged from the chimpanzees, the relation of this genus to human ancestors and whether it is a hominin is now a matter of debate. Two fossil species are described in the literature: '' A. ramidus'', which lived about 4.4 million years ago during the early Pliocene, and '' A. kadabba'', dated to approximately 5.6 million years ago (late Miocene). Initial behavioral analysis indicated that ''Ardipithecus'' could be very similar to chimpanzees; however, more recent analysis based on canine size and lack of canine sexual dimorphism indicates that ''Ardipithecus'' was characterised by reduced aggression, and that they more closely resemble bonobos. Some analyses describe ''Australopithecus'' as being sister to ''Ardipithecus ramidus'' specifically. This ...
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Paranthropus Robustus
''Paranthropus robustus'' is a species of robust australopithecine from the Early and possibly Middle Pleistocene of the Cradle of Humankind, South Africa, about 2.27 to 0.87 (or, more conservatively, 2 to 1) million years ago. It has been identified in Kromdraai, Swartkrans, Sterkfontein, Gondolin, Cooper's, and Drimolen Caves. Discovered in 1938, it was among the first early hominins described, and became the type species for the genus ''Paranthropus''. However, it has been argued by some that ''Paranthropus'' is an invalid grouping and synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus robustus''. Robust australopithecines—as opposed to gracile australopithecines—are characterised by heavily built skulls capable of producing high stresses and bite forces, as well as inflated cheek teeth (molars and premolars). Males had more heavily built skulls than females. ''P. robustus'' may have had a genetic susceptibi ...
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Orrorin
''Orrorin'' is an extinct genus of primate within Homininae from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya. The type species is ''O. tugenenesis'', named in 2001, and a second species, ''O. praegens'', assigned to the genus in 2022. Discovery and naming ''Ororrin tugenensis'' The first part of the holotype, a lower molar, was discovered by Martin Pickford in 1974 and described by Pickford (1975). The team that found the rest of the holotype of ''O. tugenensis'' was led by Brigitte Senut and Martin Pickford from the French National Museum of Natural History. Starting from 17 October 2000, 20 fossils were found at four sites in the Lukeino Formation, Kenya: of these, the fossils at Cheboit and Aragai are the oldest (), while those in Kapsomin and Kapcheberek are found in the upper levels of the formation (). ''Orrorin tugenensis'' was named and described by Senut ''et al.'' (2001). ''Orrorin praegens'' The second species, ''O. ...
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Genera
Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. '' Panthera leo'' (lion) and '' Panthera onca'' (jaguar) are two species within the genus '' Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. phylogenetic analysis should clearly demonstrate both monophyly and validity as a separate lineag ...
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Graecopithecus
''Graecopithecus'' is an extinct genus of hominid that lived in southeast Europe during the late Miocene around 7.2 million years ago. Originally identified by a single lower jawbone bearing teeth found in Pyrgos Vasilissis, Athens, Greece, in 1944, other teeth were discovered from Azmaka quarry in Bulgaria in 2012. With only little and badly preserved materials to reveal its nature, it is considered as "the most poorly known European Miocene hominoids." The creature was popularly nicknamed 'El Graeco' (word play on the Greek-Spanish painter El Greco) by scientists. In 2017, an international team of palaeontologists led by Madelaine Böhme of the Eberhard-Karls-University Tübingen, Germany, published a controversial analysis of the teeth and age of the specimens, and came to the conclusion that it could be the oldest hominin, meaning that it could be the oldest direct ancestors of humans after splitting from that of the chimpanzees. Their simultaneous study also claimed tha ...
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Australopithecus Garhi
''Australopithecus garhi'' is a species of australopithecine from the Bouri Formation in the Afar Region of Ethiopia 2.6–2.5 million years ago (mya) during the Early Pleistocene. The first remains were described in 1999 based on several skeletal elements uncovered in the three years preceding. ''A. garhi'' was originally considered to have been a direct ancestor to ''Homo'' and the human line, but is now thought to have been an offshoot. Like other australopithecines, ''A. garhi'' had a brain volume of ; a jaw which jutted out (prognathism); relatively large molars and premolars; adaptations for both walking on two legs (bipedalism) and grasping while climbing (arboreality); and it is possible that, though unclear if, males were larger than females (exhibited sexual dimorphism). One individual, presumed female based on size, may have been tall. ''A. garhi'' is the first pre-''Homo'' hominin postulated to have manufactured tools—using them in butchering—and may be counted ...
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Australopithecus Deyiremeda
''Australopithecus deyiremeda'' is an extinct species of australopithecine from Woranso–Mille, Afar Region, Ethiopia, about 3.5 to 3.3 million years ago during the Pliocene. Because it is known only from three partial jawbones, it is unclear if these specimens indeed represent a unique species or belong to the much better-known ''A. afarensis''. ''A. deyiremeda'' is distinguished by its forward-facing cheek bones and small cheek teeth compared to those of other early hominins. It is unclear if a partial foot specimen exhibiting a dextrous big toe (a characteristic unknown in any australopithecine) can be assigned to ''A. deyiremeda''. ''A. deyiremeda'' lived in a mosaic environment featuring both open grasslands and lake- or riverside forests, and anthropologist Fred Spoor suggests it may have been involved in the Kenyan Lomekwi stone tool, stone-tool industry (archaeology), industry typically assigned to ''Kenyanthropus''. ''A. deyiremeda'' coexisted with ''A. afarensis'', and ...
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