Heterokonts
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Heterokonts
The stramenopiles, also called heterokonts, are protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria. Stramenopiles are eukaryotes; most are single-celled, but some are multicellular including some large seaweeds, the brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid Heliozoa, and oomycetes. The tripartite hairs characteristic of the group have been lost in some of the included taxa – for example in most dia ...
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Ochrophyte
Ochrophytes, also known as heterokontophytes or stramenochromes, are a phylum of algae. They are the photosynthetic stramenopiles, a group of eukaryotes, organisms with a cell nucleus, characterized by the presence of two unequal flagella, one of which has tripartite hairs called mastigonemes. In particular, they are characterized by photosynthetic organelles or plastids enclosed by four membranes, with membrane-bound compartments called thylakoids organized in piles of three, chlorophyll ''a'' and ''c'' as their photosynthetic pigments, and additional pigments such as β-carotene and xanthophylls. Ochrophytes are one of the most diverse lineages of eukaryotes, containing ecologically important algae such as brown algae and diatoms. They are classified either as phylum Ochrophyta, Heterokontophyta or as subphylum Ochrophytina withing phylum Gyrista. Their plastids are of red algal origin. Description Ochrophytes are eukaryotic organisms composed of cells that are eithe ...
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Bigyromonada
Bigyromonada is a recently described non-photosynthetic lineage of stramenopiles that at present contains two classes. Description Bigyromonads are characterized by biciliate cells that feed on bacteria through phagotrophy. They are marine organisms. Taxonomy * Clade Bigyromonada Cavalier-Smith 1998 ** Class Developea Karpov & Aleoshin 2016 ex Cavalier-Smith 2017 *** Order Developayellales Doweld 2001 evelopayellida Cavalier-Smith 1987**** Family Developayellaceae Cavalier-Smith 1997 evelopayellidae***** '' Developayella'' Tong 1995 ****** '' Developayella elegans'' Tong 1995 ***** '' Develorapax'' Karpov & Aleoshin 2016 ****** '' Develorapax marinus'' Karpov & Aleoshin 2016 ** Class Pirsonea *** Order Pirsoniales ''Pirsonia'' is a non photosynthetic genus of heterokont The stramenopiles, also called heterokonts, are protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some t ... Cava ...
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Gyrista
Gyrista is a clade of stramenopile protists containing three diverse groups: the mostly photosynthetic Ochrophyta, the parasitic Pseudofungi, and the recently described group of nanoflagellates known as Bigyromonada. Members of this phylum are characterized by the presence of a helix or a double helix/ring system in the ciliary transition region. Systematics Taxonomic history Gyrista was first described in 1998 by protistologist Thomas Cavalier-Smith in his work ''A revised six-kingdom system of life'', originally as a superphylum containing two phyla: Ochrophyta, the heterokont algae; and Bigyra, which then contained the pseudofungi and bigyromonads together with the opalines. Later, the name Bigyra was modified to contain opalines, bicosoecids and labyrinthulomycetes, while the Ochrophyta, Pseudofungi and Bigyromonada remained as groups within Gyrista. Molecular phylogenetics Gyrista was seen in 2017 as the sister group to phylum Bigyra, which contains the Sag ...
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Opalozoa
Opalozoa is a subphylum of heterotrophic protists of the phylum Bigyra, and is the sister group to Sagenista. Opalozoans are non-photosynthetic heterokonts that are ancestrally phagotrophic but many times have evolved to be osmotrophic saprotrophs in the gut of vertebrate animals. Taxonomy History: phylum Opalozoa In 1993 the name “Opalozoa” referred to a group of protists that was very different from what it is now. It was a phylum composed of many unrelated zooflagellates, grouped together because of the common presence of tubular mitochondrial cristae and the lack of cortical alveoli or rigid tubular ciliary hairs ( retronemes). It also included the opalinids, proteomyxids and plasmodiophorids. Modern classification The modern taxonomy of Opalozoa, down to order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects ...
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Mesoproterozoic
The Mesoproterozoic Era is a geologic era that occurred from . The Mesoproterozoic was the first era of Earth's history for which a fairly definitive geological record survives. Continents existed during the preceding era (the Paleoproterozoic), but little is known about them. The continental masses of the Mesoproterozoic were more or less the same ones that exist today, although their arrangement on the Earth's surface was different. Major events and characteristics The major events of this era are the breakup of the Columbia supercontinent, the formation of the Rodinia supercontinent, and the evolution of sexual reproduction. This era is marked by the further development of continental plates and plate tectonics. The supercontinent of Columbia broke up between 1500 and 1350 million years ago, and the fragments reassembled into the supercontinent of Rodinia around 1100 to 900 million years ago, on the time boundary between the Mesoproterozoic and the subsequent Neoprot ...
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Protist
A protist ( ) or protoctist is any eukaryotic organism that is not an animal, land plant, or fungus. Protists do not form a natural group, or clade, but are a paraphyletic grouping of all descendants of the last eukaryotic common ancestor excluding land plants, animals, and fungi. Protists were historically regarded as a separate taxonomic kingdom known as Protista or Protoctista. With the advent of phylogenetic analysis and electron microscopy studies, the use of Protista as a formal taxon was gradually abandoned. In modern classifications, protists are spread across several eukaryotic clades called supergroups, such as Archaeplastida ( photoautotrophs that includes land plants), SAR, Obazoa (which includes fungi and animals), Amoebozoa and " Excavata". Protists represent an extremely large genetic and ecological diversity in all environments, including extreme habitats. Their diversity, larger than for all other eukaryotes, has only been discovered in rece ...
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Orth
Orth can refer to: Places * Orth, Minnesota, an unincorporated community in Nore Township, Minnesota, United States * Orth an der Donau, a town in Gänserndorf, Lower Austria, Austria * Orth House, a historic house in Winnetka, Illinois, United States * Orth C. Galloway House, a historic house in Clarendon, Arkansas, United States * Schloss Orth, a castle on Traunsee lake, Austria People * Al Orth (1872–1948), American baseball player * August Orth (1828–1901), German architect * Bertram Orth (1848–1931), German-Canadian prelate of the Catholic Church * Brodie Orth, American rugby union player * Christian Henry Orth (1773–1816), American politician * David Orth (b. 1965), Canadian actor * Eduard Orth (1902–1968), German politician * Elisa Orth (born c. 1984), Brazilian researcher * Florian Orth (b. 1989), German athlete * Frank Orth (1880–1962), American actor * Franklin Orth (1907–1970), American executive of the National Rifle Association and Preside ...
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MycoBank
MycoBank is an online database, documenting new mycological names and combinations, eventually combined with descriptions and illustrations. It is run by the Westerdijk Fungal Biodiversity Institute in Utrecht. Each novelty, after being screened by nomenclatural experts and found in accordance with the ICN (International Code of Nomenclature for algae, fungi, and plants), is allocated a unique MycoBank number before the new name has been validly published. This number then can be cited by the naming author in the publication where the new name is being introduced. Only then, this unique number becomes public in the database. By doing so, this system can help solve the problem of knowing which names have been validly published and in which year. MycoBank is linked to other important mycological databases such as Index Fungorum, Life Science Identifiers, Global Biodiversity Information Facility (GBIF) and other databases. MycoBank is one of three nomenclatural repositories reco ...
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Flagella
A flagellum (; : flagella) (Latin for 'whip' or 'scourge') is a hair-like appendage that protrudes from certain plant and animal sperm cells, from fungal spores ( zoospores), and from a wide range of microorganisms to provide motility. Many protists with flagella are known as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium '' Helicobacter pylori'', for example, uses its flagella to propel itself through the stomach to reach the mucous lining where it may colonise the epithelium and potentially cause gastritis, and ulcers – a risk factor for stomach cancer. In some swarming bacteria, the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota, the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its ...
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Kaonashia
''Kaonashia insperata'' is a species of single-celled phagotrophic stramenopiles. Described in 2023 from flagellates found in a soda lake, it is the sole species within the genus ''Kaonashia'' and family Kaonashiidae. Cells of this organism have cortical alveoli, extrusomes, a ventral groove, a short anterior flagellum covered in mastigonemes, and a longer posterior flagellum covered in thinner hairs. The cells are most similar to those found in developayellids, but a molecular phylogenetic analysis found no well-supported affinity to any specific stramenopile group. Etymology The genus ''Kaonashia'' is named after the fictional ghost from the Hayao Miyazaki film ''Spirited Away''. The specific epithet ''insperata'' () references the unanticipated phylogenetic placement of this organism given its appearance. Description Cellular organization ''Kaonashia insperata'' is a species of free-living phagotrophic flagellates. The cells are roughly ovoid or bean-shaped, 7–13 μm ...
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SAR Supergroup
SAR is a highly diverse clade of eukaryotes, often considered a supergroup, that includes stramenopiles (heterokonts), alveolates, and rhizarians. It is a node-based taxon (under the Sar name), including all descendants of the three groups' last common ancestor, and comprises most of the now-rejected Chromalveolata. Their sister group has been found to be telonemids, with which they make up the TSAR clade. Harosa is sometimes used synonymously with TSAR. Etymology The name SAR is an acronym derived from the first letters of its three constituent clades; it has been alternatively spelled RAS. The term Harosa (at the subkingdom level) has also been used, with Stramenopiles replaced by its synonym Heterokonta in this variant of the acronym. History of discovery Before the discovery of the SAR supergroup, stramenopiles and alveolates were classified in the supergroup Chromalveolata alongside haptophytes and cryptomonads, being believed to have acquired plastids th ...
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Supergroup (biology)
A supergroup or super-group, in systematics, is a large group of organisms that monophyletic, share one common ancestor and have important defining characteristics. It is an informal, mostly arbitrary rank in biology, biological Taxonomy (biology), taxonomy that is often greater than phylum or kingdom (biology), kingdom, although some supergroups are also treated as phylum, phyla. Eukaryotic supergroups Since the decade of the 2000s, the Eukaryote#Phylogeny, eukaryotic tree of life (abbreviated as eToL) has been divided into 5–8 major groupings called 'supergroups'. These groupings were established after the idea that only monophyletic groups should be accepted as ranks, as an alternative to the use of paraphyletic kingdom Protista. In the early days of the eToL six traditional supergroups were considered: Amoebozoa, Opisthokonta, "Excavata", Archaeplastida, "Chromalveolata" and Rhizaria. Since then, the eToL has been rearranged profoundly, and most of these groups were found as pa ...
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