Elephantimorpha
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Elephantimorpha
Elephantimorpha is a clade of proboscideans that contains the Mammutidae (mastodons), as well as Elephantida ( amebelodonts, choerolophodonts, gomphotheres, stegodontids and elephantids). All members of Elephantimorpha have the horizontal tooth replacement typical of modern elephants, where the cheek teeth progressively migrate forwards in the jaw like a conveyor belt. This a distinctive characteristic of this group (synapomorphy) and is not found in more primitive non-elephantimorph Elephantiformes. Members of Elephantida are distinguished from mammutids by the reduced angular process of the lower jaw. Like modern elephants, the ancestor of Elephantimorpha was likely capable of communicating via infrasonic calls. While early elephantimorphs generally had lower jaws where the fused front part (the mandibular symphysis) was greatly elongated with well developed lower tusks/incisors, from the Late Miocene onwards, many groups convergently developed brevirostrine (shortened) lowe ...
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Proboscideans
Proboscidea (; , ) is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three living species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. Extinct members of Proboscidea include the deinotheres, mastodons, gomphotheres and stegodonts. The family Elephantidae also contains several extinct groups, including mammoths and ''Palaeoloxodon''. Proboscideans include some of the largest known land mammals, with the elephant '' Palaeoloxodon namadicus'' and mastodon ''"Mammut" borsoni'' suggested to have body masses surpassing , rivalling or exceeding paraceratheres (the otherwise largest known land mammals) in size. The largest extant proboscidean is the African bush elephant, with a world record of size of at the shoulder and . In addition to their enor ...
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Gomphotheriidae
Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as '' Gomphotherium'' had long lower jaws with tusks, the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks and outlasted the long-jawed gomphotheres. This change made them look very similar to modern elephants, an example of parallel evolution. During the Pliocene and Early Pleistocene, the diversity of gomphotheres declined, ultimately becoming extinct outside of the Americas. The last two genera, '' Cuvieronius'' ranging southern North America to w ...
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Gomphotheres
Gomphotheres are an extinct group of proboscideans related to modern Elephant, elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a Paraphyly, paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as ''Gomphotherium'' had long lower jaws with tusks, the Plesiomorphy and symplesiomorphy, ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks and outlasted the long-jawed gomphotheres. This change made them look very similar to modern elephants, an example of parallel evolution. During the Pliocene and Early Pleistocene, the diversity of gomphotheres declined, ultimately becoming extinct outside of the Americas. The last two genera, ''Cuv ...
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Gomphothere
Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as '' Gomphotherium'' had long lower jaws with tusks, the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks and outlasted the long-jawed gomphotheres. This change made them look very similar to modern elephants, an example of parallel evolution. During the Pliocene and Early Pleistocene, the diversity of gomphotheres declined, ultimately becoming extinct outside of the Americas. The last two genera, '' Cuvieronius'' ranging southern North America to weste ...
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Mammutidae
Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons (genus ''Mammut''), which inhabited North America from the Late Miocene (around 8 million years ago) until their extinction at the beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, "nipple" and "tooth", referring to their characteristic teeth. Description Mammutids are characterised by their zygodont molars, where pairs of parallel cusps are merged into sharp-sided riges, which are morphologically conservative and differ little between mammutid species.'''' Like other members of Elephantimorpha, mammutids exhibited horizontal tooth replacement like modern elephants. Some authors have argued that horizontal tooth replacement evolved in parallel in mammu ...
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Gomphotherium Annectens
''Gomphotherium'' (; "nail beast" for its double set of straight tusks) is an extinct genus of gomphothere proboscidean from the Neogene of Eurasia, Africa and North America. It is the most diverse genus of gompothere, with over a dozen valid species. The genus is probably paraphyletic. Description Most species of ''Gomphotherium'' were similar in size to the Asian elephant, with ''G. productum'' (known from a 35-year-old male) measuring tall and weighing . The largest species ''G. steinheimense'', known from a complete 37-year-old male found in Mühldorf, Germany, measured up to tall and weighed . ''Gomphotherium'', like most basal elephantimorphs, had an elongated lower jaw which bore tusks. Species of ''Gomphotherium'' are defined by their conservative molar morphology, which includes "trilophed intermediate molars, third molars with three to four loph(id)s, and pretrite half-loph(id)s typically with anterior and posterior accessory conules that form trefoil-patterne ...
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Elephantiformes
Elephantiformes is a suborder within the order Proboscidea. Members of this group are primitively characterised by the possession of upper tusks, an elongated mandibular symphysis (the frontmost part of the lower jaw) and lower tusks, and the retraction of the facial region of the skull indicative of the development of a trunk. The earliest known member of the group, '' Dagbatitherium'' is known from the Eocene (Lutetian) of Togo, which is only known from isolated teeth, while other primitive elephantiforms like '' Phiomia'' and '' Palaeomastodon'' are known from the Early Oligocene onwards. ''Phiomia'' and ''Palaeomastodon'' are often collectively referred to as "palaeomastodonts" and assigned to the family Palaeomastodontidae. Most diversity of the group is placed in the subclade Elephantimorpha, which includes mastodons (family Mammutidae), as well as modern elephants and gomphotheres (Elephantida), which are distinguished from more primitive elephantiforms by the development ...
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Eubelodon
''Eubelodon'' is an extinct genus of gomphothere (a family in the order Proboscidea, which also includes modern elephants) which lived in North America during the Miocene Epoch. It contains a single species: ''Eubelodon morrilli''. Description ''Eubelodon'' is considered to be a trilophodont gomphothere. It has the highly unusual combination of retaining a long lower jaw, but having lost the lower tusks, a combination only shared with fellow North American gomphothere ''Gnathabelodon'' and the choerolophodontid '' Choerolophodon.'' Fossil distribution Fossils are restricted to what is now the Great Plains of the United States. Remains were found in the Poison Ivy Quarry, Antelope, Brown County, Nebraska, and Tripp County, South Dakota. Taxonomy ''Eubelodon'' was named by Erwin Hinckly Barbour in 1914. It was synonymized subjectively with ''Trilophodon'' by Osborn in 1918 and again by Tobien in 1973 with '' Gomphotherium''. It was assigned to Gomphotheriidae by Erwin ...
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Stegodontidae
Stegodontidae is an extinct family of proboscideans from Africa and Asia (with a single occurrence in Europe) from the Early Miocene (at least 17.3 million years ago) to the Late Pleistocene. It contains two genera, the earlier ''Stegolophodon'', known from the Miocene of Asia and the later '' Stegodon,'' from the Late Miocene to Late Pleistocene of Africa and Asia (with a single occurrence in Greece) which is thought to have evolved from the former. The group is noted for their plate-like lophs on their teeth, which are similar to elephants and different from those of other extinct proboscideans like gomphotheres Gomphotheres are an extinct group of proboscideans related to modern Elephant, elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Ple ... and mammutids, with both groups having a proal jaw movement utilizing forward strokes of the lower jaw. These simila ...
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Anancus
''Anancus'' is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago. Taxonomy ''Anancus'' was named by Auguste Aymard in 1855. It is traditionally allocated to Gomphotheriidae, often as the only member of the subfamily Anancinae. Recently, some authors have excluded ''Anancus'' along with other tetralophodont gomphotheres from Gomphotheriidae, and regarded them as members of Elephantoidea instead. Description Two largely complete individuals of ''Anancus arvernensis'' reached shoulder heights of around , with a volumetric estimate suggesting a body mass of around , comparable to living African bush elephants. The tusks were largely straight and lacked enamel (though enamel was present in juvenilesTheodorou, G., Spjeldnaes, N., Hanken, N. M., Lauritzen, S. E., Velitzelos, E., Athanassiou, A., et al. (2000). Des ...
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