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Candelariella Immarginata
''Candelariella immarginata'' is a species of parasitic, saxicolous (rock-dwelling) lichen in the family Candelariaceae. Found in the United States, it was formally described as a new species in 2007 by Swedish lichenologist Martin Westberg. The type specimen was collected in the desert west of Grantsville, Utah at an elevation of ; here it was found growing on dry exposed quartzite. At the time of publication, it had also been found in another location in Utah, and two locations in Nevada; its general geographic range is the Great Basin in western North America. In 2015, it was recorded from the White Mountains of California. The species is lichenicolous on ''Aspicilia'' species and pyrenocarpous lichens. The specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, ...
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Parasitism
Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of life. The entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as the agents of malaria, sleeping sickness, and amoebic dysentery; animals such as hookworms, lice, mosquitoes, and vampire bats; fungi such as Armillaria mellea, honey fungus and the agents of ringworm; and plants such as mistletoe, dodder, and the Orobanchaceae, broomrapes. There are six major parasitic Behavioral ecology#Evolutionarily stable strategy, strategies of exploitation of animal hosts, namely parasitic castration, directly transmitted parasitism (by contact), wikt:trophic, trophicallytransmitted parasitism (by being eaten), Disease vector, vector-transmitted paras ...
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Botanical Name
A botanical name is a formal scientific name conforming to the ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the '' International Code of Nomenclature for Cultivated Plants'' (ICNCP). The code of nomenclature covers "all organisms traditionally treated as algae, fungi, or plants, whether fossil or non-fossil, including blue-green algae ( Cyanobacteria), chytrids, oomycetes, slime moulds and photosynthetic protists with their taxonomically related non-photosynthetic groups (but excluding Microsporidia)." The purpose of a formal name is to have a single name that is accepted and used worldwide for a particular plant or plant group. For example, the botanical name '' Bellis perennis'' denotes a plant species which is native to most of the countries of Europe and the Middle East, where it has accumulated various names in many languages. Later, the plant w ...
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Lichens Of The Southwestern United States
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Candelariella
''Candelariella'' is a genus of bright yellow, ocher, or greenish yellow crustose or squamulose lichens in the family Candelariaceae. Members of the genus are commonly called eggyolk lichens, goldspeck lichens, or yolk lichens. The genus was circumscribed in 1894 by Swiss lichenologist Johannes Müller Argoviensis, with ''Candelariella vitellina'' assigned as the type species. Characteristics The key feature of ''Candelariella'' species are the distinct yellow apothecia. Although all species are very small, even the smallest can be identified by the lemon-yellow to orange-yellow discs. Most species have a yellow thallus, although ''Candelariella antennaria'' is one example with a grey thallus. Some species are pycnidiate. This genus will generally have all spot tests emerge as negative, although K tests may have an orange or reddish colour on some species' apothecia. Spore count between species varies from 8 to 32 simply or thinly septate spores. Spores often hold one to tw ...
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Pulvinic Acid
Pulvinic acids are natural chemical pigments found in some lichens, derived biosynthetically from the aromatic amino acids phenylalanine and tyrosine, via dimerization and oxidative ring-cleavage of arylpyruvic acids, a process that also produces the related pulvinones. Hydroxypulvinic acid pigments (pulvinic acid type family of pigments) have been found in ''Boletus'' (e.g. '' Boletus erythropus''), '' Boletinus'', '' Chalciporus'', '' Gyrodon'', '' Leccinum'', '' Pulveroboletus'', '' Suillus'' (e.g. ''Suillus luteus'', '' Suillus bovinus'', and '' Suillus grevillei''), ''Paxillus'' (e.g. '' Paxillus involutus''), '' Serpula'' (e.g. '' Serpula lacrymans''), ''Xerocomus'' (e.g. '' Xerocomus chrysenteron''), '' Hygrophoropsis'' (e.g. '' Hygrophoropsis aurantiaca''), '' Retiboletus'' (e.g. '' Retiboletus nigerrimus''), '' Pulveroboletus'' (e.g. '' Pulveroboletus auriflammeus''), and are generally characteristic of Boletales. In addition to pulvinone, derivatives and related pigm ...
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Vulpinic Acid
Vulpinic acid is a natural product first found in and important in the symbiosis underlying the biology of lichens. It is a simple methyl ester derivative of its parent compound, pulvinic acid, and a close relative of pulvinone, both of which derive from aromatic amino acids such as phenylalanine via secondary metabolism. The roles of vulpinic acid are not fully established, but may include properties that make it an antifeedant for herbivores. The compound is relatively toxic to mammals. Chemical description Vulpinic acid was first isolated from lichens in 1925. As an isolated, purified substance, it is bright yellow in color. Vulpinic acid is derived biosynthetically by esterification from pulvinic acid; pulvinate itself derives from the aromatic amino acids phenylalanine and tyrosine, via dimerization and oxidative ring-cleavage of arylpyruvic acids, a process that also produces the related pulvinones. There have been several chemical syntheses reported for vulpinic ac ...
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High-performance Liquid Chromatography
High-performance liquid chromatography (HPLC), formerly referred to as high-pressure liquid chromatography, is a technique in analytical chemistry used to separate, identify, and quantify each component in a mixture. It relies on pumps to pass a pressurized liquid solvent containing the sample mixture through a column filled with a solid adsorbent material. Each component in the sample interacts slightly differently with the adsorbent material, causing different flow rates for the different components and leading to the separation of the components as they flow out of the column. HPLC has been used for manufacturing (''e.g.'', during the production process of pharmaceutical and biological products), legal (''e.g.'', detecting performance enhancement drugs in urine), research (''e.g.'', separating the components of a complex biological sample, or of similar synthetic chemicals from each other), and medical (''e.g.'', detecting vitamin D levels in blood serum) purposes. Chrom ...
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Secondary Metabolite
Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the normal growth, development, or reproduction of the organism. Instead, they generally mediate ecological interactions, which may produce a selective advantage for the organism by increasing its survivability or fecundity. Specific secondary metabolites are often restricted to a narrow set of species within a phylogenetic group. Secondary metabolites often play an important role in plant defense against herbivory and other interspecies defenses. Humans use secondary metabolites as medicines, flavourings, pigments, and recreational drugs. The term secondary metabolite was first coined by Albrecht Kossel, a 1910 Nobel Prize laureate for medicine and physiology in 1910. 30 years later a Polish botanist Friedrich Czapek described secondary ...
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