Brachiopod Taxonomy
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Brachiopod Taxonomy
Brachiopods (), phylum (biology), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachio ...
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Lingula Anatina
''Lingula anatina'' is a brachiopod species in the genus '' Lingula''. Like others in its genus, ''L. anatina'' is a filter feeder Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a s ... that uses a lophophore to extract food from water. They burrow in the sand of their brackish intertidal habitat. References Lingulata Animals described in 1801 {{brachiopod-stub ...
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Yolk Sac
The yolk sac is a membranous wikt:sac, sac attached to an embryo, formed by cells of the hypoblast layer of the bilaminar embryonic disc. This is alternatively called the umbilical vesicle by the Terminologia Embryologica (TE), though ''yolk sac'' is far more widely used. The yolk sac is one of the fetal membranes and is important in early embryonic blood supply. In humans much of it is incorporated into the primordial gut (anatomy), gut during the fourth week of embryonic development. In humans The yolk sac is the first element seen within the gestational sac during pregnancy, usually at three days gestation. The yolk sac is situated on the front (ventral) part of the embryo; it is lined by extra-embryonic endoderm, outside of which is a layer of extra-embryonic mesenchyme, derived from the epiblast. Blood is conveyed to the wall of the yolk sac by the primitive aorta and after circulating through a wide-meshed capillary plexus, is returned by the vitelline veins to the tubul ...
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Capitanian Mass Extinction Event
The Capitanian mass extinction event (also known as the end-Guadalupian extinction event, the Guadalupian-Lopingian boundary mass extinction, the pre-Lopingian crisis, or the Middle Permian extinction) was a major mass extinction event that occurred towards the end of the Capitanian age and Guadalupian (Middle Permian) epoch of the Permian period. The mass extinction occurred during a period of decreased species richness and increased extinction rates. It is often called the end-Guadalupian extinction event because of its initial recognition between the Guadalupian and Lopingian series; however, more refined stratigraphic study suggests that extinction peaks in many taxonomic groups occurred within the Guadalupian, in the latter half of the Capitanian age.Bond, D. P. G., Wignall, P. B., Wang, W., Izon, G., Jiang, H. S., Lai, X. L., Sund, Y.-D., Newtona, R.J., Shaoe, L.-Y., Védrinea, S. & Cope, H. (2010). "The mid-Capitanian (Middle Permian) mass extinction and carbon isotope reco ...
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Paleozoic
The Paleozoic ( , , ; or Palaeozoic) Era is the first of three Era (geology), geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at the start of the Mesozoic Era. The Paleozoic is subdivided into six period (geology), geologic periods (from oldest to youngest), Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian. The name ''Paleozoic'' was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (geologist), John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Ancient Greek, Greek ''palaiós'' (π� ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Tommotiid
Tommotiids are an extinct group of Cambrian invertebrates thought to be early total-group lophophorates (the group containing Bryozoa, Brachiopoda, and Phoronida), including members of the lophophorate stem group, as well as early diverging members of the crown group. The majority of tommotiids are mineralised with calcium phosphate rather than calcium carbonate, although silicified examples hint that some species bore carbonate or carbonaceous sclerites. '' Micrina'' and '' Paterimitra'' possess bivalved shells in their larval phases, which preserve characters that might position them in the Linguliformea and Rhynchonelliformea stem lineages respectively. This would indicate that the brachiopod shell represents the retention of a larval character. For a long part of their history, the tommotiids were only known from disarticulated shells - a complete organism had not been found. The 2008 discovery of '' Eccentrotheca'' offered the first insight into a complete organism, a ...
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Phoronid
Phoronids (Taxonomy (biology), taxonomic name Phoronida, sometimes called horseshoe worms) are a small phylum of marine animals that Filter feeder, filter-feed with a lophophore (a "crown" of tentacles), and build upright tubes of chitin to support and protect their soft bodies. They live in most of the oceans and seas, including the Arctic Ocean but excluding the Antarctic Ocean, and between the intertidal zone and about 400 meters down. Most adult phoronids are 2 cm long and about 1.5 mm wide, although the largest are 50 cm long. The name of the group comes from its type genus: ''Phoronis#Etymology, Phoronis''. Overview The bottom end of the body is an ampulla (a flask-like swelling), which anchors the animal in the tube and enables it to retract its body very quickly when threatened. When the lophophore is extended at the top of the body, cilia (little hairs) on the sides of the tentacles draw food particles to the mouth, which is inside and slightly to o ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic animal, aquatic invertebrate animals, nearly all living in sedentary Colony (biology), colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeder, filter feeding. Most Marine (ocean), marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the Stenolaemata, marine bryozoans (Stenolaemata), Phylactolaemata, freshwater bryozoans (Phylactolaemata), and Gymnolaemata, mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. Originally all of the crown group Bryozoa were colonial, but as an adaptation to a mesopsammal (interstitial spaces in marine sand) life or to deep-sea habitats, secondarily solitary forms have since evolved. Solitary species have been described i ...
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Lophophorata
The Lophophorata (also called Tentaculata; not to be confused with Tentaculata Eschscholtz 1825, a class within the Ctenophora) are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic animal, aquatic invertebrate animals, nearly all living in sedentary Colony (biology), colonies. Typically about long, they have a spe .... They have a lophophore. Molecular phylogenetic analyses suggest that lophophorates are protostomes, but on morphological grounds they have been assessed as deuterostomes. Fossil finds of the " tommotiid" '' Wufengella'' suggest that they evolved from worm-like animals that resembled annelids. References {{Protostome-stub ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is Cladogram#Homoplasies, homoplasy. The recurrent evolution of flight is a classic example, as flying pterygota, insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas ''homology (biology), homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different trai ...
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Terebratulida
Terebratulids are one of only three living orders of articulate brachiopods, the others being the Rhynchonellida and the Thecideida. Craniida and Lingulida include living brachiopods, but are inarticulates. The name, Terebratula, may be derived from the Latin "terebra", meaning "hole-borer". The perceived resemblance of terebratulid shells to ancient Roman oil lamps gave the brachiopods their common name "lamp shell". Terebratulids typically have biconvex shells that are usually ovoid to circular in outline. They can be either smooth or have radial ribbing. The lophophore support is loop shaped in contrast to the spiralia of similar looking spiriferids. Terebratulids are also distinguished by a very short hinge line, and the shell is punctate in microstructure. There is a circular pedicle opening, or foramen, located in the beak. Terebratulids may have evolved from Atrypids during the early or Middle Silurian. Early genera were almost circular to elongate-oval, with s ...
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Class (biology)
In biological classification, class () is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are domain, kingdom, phylum, order, family, genus, and species, with class ranking between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name – and not just called a ''top-level genus'' ''(genus summum)'' – was first introduced by French botanist Joseph Pitton de Tournefort in the classification of plants that appeared in his '' Eléments de botanique'' of 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, whic ...
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