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Barbourula
''Barbourula'' is a genus of amphibian commonly referred to as jungle toads. They are small toads of the fire-bellied toad family, Bombinatoridae, found in the Philippines and Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda .... Species References Amphibian Species of the World, ''Barbourula''Living Underworld, Bombinatoridae
Amphibian genera
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Barbourula Busuangensis
''Barbourula busuangensis'' is a species of frog in the family Bombinatoridae. It is also known under the common names Philippine flat-headed frog, Palawan flat-headed frog, Busuanga jungle toad, Busuanga disk-tongued toad, and Philippine aquatic frog. It is endemic to the Busuanga, Culion, Balabac, and Palawan islands in the Philippines. It is an inhabitant of clear-water streams and is threatened by habitat loss. Description ''Barbourula busuangensis'' is a large aquatic species of frog. Its body is flattened dorso-ventrally and its nostrils and eyes are located at the top of its head. It has no external tympani. Both fore and hind limbs are robust and powerful, and the digits of the hands are webbed as well as those of the feet. The surface of the body is covered with fine tubercles. This frog is a dark greenish-black in colour with some faint pale green hatching and spotting on the hind limbs. Distribution and habitat This frog is endemic to the western Philippines where i ...
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Bornean Flat-headed Frog
The Bornean flat-headed frog (''Barbourula kalimantanensis'') is a species of frog in the family Bombinatoridae. For many years, it was thought to be the only frog with no lungs. However, micro CT scanning revealed that, like all other known frog species, the Bornean flat-headed frog has lungs, though they are tiny. Djoko Iskandar, an Indonesian zoologist, first described the Bornean flat-headed frog based on a single specimen. However, he did not think the frog was lungless until he was in the field with David Bickford's expedition. Bickford ''et al.'' dissected some of the specimens and were unable to see the lungs, leading them to incorrectly conclude that no lungs were present. Description This frog grows to a snout-to-vent length of for females and males are slightly smaller. It has a dorsoventrally flattened body with a broad head with a rounded snout. The limbs are robust and both hands and feet are fully webbed. The dorsal skin is smooth with small tubercles on the pos ...
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Barbourula Busuangensis05
''Barbourula'' is a genus of amphibian commonly referred to as jungle toads. They are small toads of the fire-bellied toad family, Bombinatoridae, found in the Philippines and Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda .... Species References Amphibian Species of the World, ''Barbourula''Living Underworld, Bombinatoridae
Amphibian genera
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Bombinatoridae
Bombinatoridae is a family (biology), family of toads found in Eurasia. Species of the family have flattened bodies and some are highly toxic. Taxonomy and systematics Fossil specimens of the genus ''Bombina'' are known from the Pliocene to the Pleistocene. The earliest fossil specimens are ''Eobarbourula'' from the Eocene of India, and ''Hatzegobatrachus'' from Late Cretaceous of Hateg island, Romania. The genus ''Barbourula'' was considered to be situated intermediate between ''Discoglossus'' and ''Bombina'', but closer to the latter, so was added to the Bombinatoridae when that family was split from the Discoglossidae. Genera Currently, there are two extant and at least two extinct genera recognised in the family Bombinatoridae:Bombinatoridae
at the American Museum of Natural History's Amphibian Species of the World w ...
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Edward Harrison Taylor
Edward Harrison Taylor (April 23, 1889 – June 16, 1978) was an American herpetologist from Missouri. Early life Taylor was born in Maysville, Missouri, to George and Loretta Taylor. He had an older brother, Eugene. Taylor studied at the University of Kansas in Lawrence, Kansas, graduating with a B.A. in 1912. Field trips during his time at the University of Kansas with Dr. Clarence McClung and Dr. Roy Moody helped prepare Taylor for his future endeavors. Between 1916 and 1920 he returned briefly to Kansas to finish his M.A. Career Upon completing his bachelor's degree, Taylor went to the Philippines, where at first he held a teacher's post in a village in central Mindanao, in the Industrial School for Manobo now known as the Agusan del Sur State College of Agriculture and Technology (ASSCAT). He collected and studied the local herpetofauna extensively and published many papers. He returned to the Philippines after completing his master's degree and was appointed Chief of ...
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Gladwyn Kingsley Noble
Gladwyn Kingsley Noble (September 20, 1894 – December 9, 1940) was an American zoologist who served as the head curator for the department of herpetology and the department of experimental biology at the American Museum of Natural History. Noble received bachelor's and master's degrees from Harvard University in 1917 and 1918, respectively, and a Ph.D. from Columbia University in 1922. He joined the herpetology department in 1922 as a research assistant and assistant curator in 1917, and became the chairman of the department in 1924. He later formed the Department of Experimental Biology in 1928, and served as the chairman of both departments until his death on December 9, 1940, from a streptococcal throat infection. Background Noble's father was Gilbert Clifford Noble, one of the founders of what would become Barnes & Noble bookstores and publishing house. Gilbert Clifford Noble joined the Arthur Hinds & Company firm in 1886 after graduating from Harvard College. In 1894, he wa ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Amphibian
Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant taxon, extant (living) amphibians belong to the monophyletic subclass (biology), subclass Lissamphibia, with three living order (biology), orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater ecosystem, freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their biological life cycle, life cycle typically starts out as aquatic animal, aquatic larvae with gills known as tadpoles, but some species have devel ...
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Philippines
The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a total area of roughly 300,000 square kilometers, which are broadly categorized in Island groups of the Philippines, three main geographical divisions from north to south: Luzon, Visayas, and Mindanao. With a population of over 110 million, it is the world's List of countries and dependencies by population, twelfth-most-populous country. The Philippines is bounded by the South China Sea to the west, the Philippine Sea to the east, and the Celebes Sea to the south. It shares maritime borders with Taiwan to the north, Japan to the northeast, Palau to the east and southeast, Indonesia to the south, Malaysia to the southwest, Vietnam to the west, and China to the northwest. It has Ethnic groups in the Philippines, diverse ethnicities and Culture o ...
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Borneo
Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda Islands, located north of Java Island, Java, west of Sulawesi, and east of Sumatra. The island is crossed by the equator, which divides it roughly in half. The list of divided islands, island is politically divided among three states. The sovereign state of Brunei in the north makes up 1% of the territory. Approximately 73% of Borneo is Indonesian territory, and in the north, the East Malaysian states of Sabah and Sarawak make up about 26% of the island. The Malaysian federal territory of Labuan is situated on a small island just off the coast of Borneo. Etymology When the sixteenth-century Portuguese explorer Jorge de Menezes made contact with the indigenous people of Borneo, they referred to their island as ''Pulu K'lemantang'', which ...
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