Annelid Anatomy
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Annelid Anatomy
The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The annelids are Symmetry in biology, bilaterally symmetrical, Triploblasty, triploblastic, coelomate, invertebrate organisms. They also have Parapodium, parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistics, Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Siboglinidae, Pogonophora, Echiura and Sipuncula, previ ...
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Ediacaran
The Ediacaran ( ) is a geological period of the Neoproterozoic geologic era, Era that spans 96 million years from the end of the Cryogenian Period at 635 Million years ago, Mya to the beginning of the Cambrian Period at 538.8 Mya. It is the last period of the Proterozoic geologic eon, Eon as well as the last of the so-called "Precambrian supereon", before the beginning of the subsequent Cambrian Period marks the start of the Phanerozoic Eon, where recognizable fossil evidence of life becomes common. The Ediacaran Period is named after the Ediacara Hills of South Australia, where trace fossils of a diverse community of previously unrecognized lifeforms (later named the Ediacaran biota) were first discovered by geologist Reg Sprigg in 1946. Its status as an official geological period was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period took namesake from the Ediacara Hills ...
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Siboglinidae
Siboglinidae is a family (biology), family of polychaete Annelida, annelid worms whose members made up the former phylum, phyla Pogonophora and Vestimentifera (the giant tube worms). The family is composed of around 100 species of vermiform creatures which live in thin tubes buried in sediment (Pogonophora) or in tubes attached to hard substratum (Vestimentifera) at ocean depths ranging from . They can also be found in association with hydrothermal vents, methane seeps, sunken plant material, and Whale fall, whale carcasses. The first specimen was dredged from the waters of Indonesia in 1900. These specimens were given to French zoologist Maurice Caullery, who studied them for nearly 50 years. Anatomy Most siboglinids are less than in diameter, but in length. They inhabit tubular structures composed of chitin which are fixed to rocks or substrates. The tubes are often clustered together in large colonies. Their bodies are divided into four regions. The anterior end is called th ...
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Hirudinea
Leeches are segmented parasitic or predatory worms that comprise the subclass Hirudinea within the phylum Annelida. They are closely related to the oligochaetes, which include the earthworm, and like them have soft, muscular segmented bodies that can lengthen and contract. Both groups are hermaphrodites and have a clitellum, but leeches typically differ from the oligochaetes in having suckers at both ends and ring markings that do not correspond with their internal segmentation. The body is muscular and relatively solid; the coelom, the spacious body cavity found in other annelids, is reduced to small channels. The majority of leeches live in freshwater habitats, while some species can be found in terrestrial or marine environments. The best-known species, such as the medicinal leech, ''Hirudo medicinalis'', are hematophagous, attaching themselves to a host with a sucker and feeding on blood, having first secreted the peptide hirudin to prevent the blood from clotting ...
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Branchiobdellida
Branchiobdellida is an order of freshwater leech-like clitellates that are obligate ectosymbionts or ectoparasites, mostly of astacoidean crayfish. They are found in the Northern Hemisphere and have a holarctic distribution in East Asia, the Euro-Mediterranean region and North and Central America, with the greatest species diversity being in North and Central America. Taxonomy The exact relationship of this group of clitellates with other groups has been the subject of debate for a century or more. The American zoologist Perry C. Holt wrote his doctoral thesis on branchiobdellids and devoted his 45 years of research to the taxon. With his colleague, the American zoologist Richard L. Hoffman, he described 8 new genera and 75 new species, and re-described various other species. In 1965, he came to the conclusion that the group should be raised from a family to an order, a sister group to the oligochaetes and leeches. Forty years later, molecular data from rDNA and mitochond ...
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Oligochaeta
Oligochaeta () is a subclass of soft-bodied animals in the phylum Annelida, which is made up of many types of aquatic and terrestrial worms, including all of the various earthworms. Specifically, oligochaetes comprise the terrestrial megadrile earthworms (some of which are semiaquatic or fully aquatic), and freshwater or semiterrestrial microdrile forms, including the tubificids, pot worms and ice worms ( Enchytraeidae), blackworms ( Lumbriculidae) and several interstitial marine worms. With around 10,000 known species, the Oligochaeta make up about half of the phylum Annelida. These worms usually have few setae (chaetae) or "bristles" on their outer body surfaces, and lack parapodia, unlike polychaeta. Diversity Oligochaetes are well-segmented worms and most have a spacious body cavity (coelom) used as a hydroskeleton. They range in length from less than up to in the 'giant' species such as the giant Gippsland earthworm (''Megascolides australis'') and the Mekong ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Clitellata
The Clitellata are a class of annelid worms, characterized by having a clitellum – the 'collar' that forms a reproductive cocoon during part of their life cycles. The clitellates comprise around 8,000 species. Unlike the class of Polychaeta, they do not have parapodia and their heads are less developed. Characteristics Clitellate annelids are segmented worms characterised by the clitellum or girdle which is located near the head end of mature individuals. The mouth is on the ventral surface and is overhung by the prostomium (proboscis). The brain is not located in the head but in one of the body segments. The clitellum is formed by a modification of several segments, and either includes the female gonopores or is located just behind them. During copulation, this glandular tissue secretes mucus that keeps the paired individuals together while they exchange sperm. Afterwards it secretes material that forms a cocoon that encircles the animal's body and encloses the eggs and ...
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Maldanomorpha
Maldanomorpha is a monophyletic group, or clade, of polychaete worms in the phylum Annelida The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to variou .... Several phylogenetic analyses based on morphological and molecular data have shown that this clade unites the families Arenicolidae and Maldanidae. The two main synapomorphies, morphological characters described to unite both families, are a lecithotrophic larval development and the presence of an uncinus, a hook-like structure with barbs. References Polychaetes Animals described in 1995 Annelid unranked clades {{Annelid-stub ...
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Terebelliformia
Terebelliformia is a group of polychaete worms in the phylum Annelida The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to variou .... Phylogeny The internal relationships within the clade Terebelliformia are the following: References Polychaetes {{Annelid-stub ...
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Capitellidae
Capitellidae is a polychaete worm family in the subclass Scolecida Scolecida is an Infraclass (biology), infraclass of polychaete worms. Scolecids are mostly unselective Detritivore, deposit feeders on marine detritus. Characteristics Scolecids have Parapodium, parapodia with rami that are all alike. The pros .... Genera * '' Abyssocapitella'' * '' Amastigos'' * '' Anotomastus'' * '' Baldia'' * '' Barantolla'' * '' Branchiocapitella'' * '' Capitella'' * '' Capitellethus'' * '' Capitobranchus'' * '' Dasybranchetus'' * '' Dasybranchus'' * '' Decamastus'' * '' Ditrocha'' * '' Dodecaseta'' * '' Eunotomastus'' * '' Heteromastides'' * '' Heteromastus'' * '' Leiocapitella'' * '' Leiocapitellides'' * '' Leiochrides'' * '' Leiochrus'' * '' Lumbricomastus'' * '' Mastobranchus'' * '' Mediomastus'' * '' Neoheteromastus'' * '' Neomediomastus'' * '' Neonotomastus'' * '' Neopseudocapitella'' * '' Nonatus'' * '' Notodasus'' * '' Notomastus'' * '' Octocapitella'' * '' Paracapitella'' * '' ...
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Echiura
The Echiura, or spoon worms, are a small group of ocean, marine animals. Once treated as a separate phylum, they are now considered to belong to Annelida. Annelids typically have their bodies divided into Segmentation (biology), segments, but echiurans have secondarily lost their segmentation. The majority of echiurans live in burrows in soft sediment in shallow water, but some live in rock crevices or under boulders, and there are also deep sea forms. More than 230 species have been described. Spoon worms are cylindrical, soft-bodied animals usually possessing a non-retractable proboscis which can be rolled into a scoop-shape to feed. In some species the proboscis is ribbon-like, longer than the trunk and may have a forked tip. Spoon worms vary in size from less than a centimetre in length to more than a metre. Most are deposit feeders, collecting detritus from the sea floor. Fossils of these worms are seldom found and the earliest known fossil specimen is from the Darriwilian, ...
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