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Bryoria Forsteri
''Bryoria'' is a genus of lichenized fungi in the family Parmeliaceae. Many members of this genus are known as horsehair lichens. The genus has a widespread distribution, especially in boreal and cool temperate areas. Taxonomy ''Bryoria'' was circumscribed in 1977 by lichenologists Irwin Brodo and David Hawksworth, with ''Bryoria trichodes'' as the type species. Molecular studies have revealed that the genus originated approximately 11.5 million years ago during the Miocene period. The genus comprises several sections, including a recently revised understanding of section ''Bryoria'', which is now known to be polyphyletic and divided into two distinct clades. The first group (''Bryoria'' clade 1) has been placed in an emended section ''Americanae'', which is primarily restricted to western North America and the Himalayan region, with ''B. americana'' being the only widely distributed species in this section. Molecular studies by Velmala et al. (2014) and Boluda et al. ...
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Bryoria Nadvornikiana
''Bryoria nadvornikiana'', commonly known as the spiny grey horsehair lichen or the blonde horsehair lichen, is a species of horsehair lichen in the family Parmeliaceae The Parmeliaceae is a large and diverse family of Lecanoromycetes. With over 2700 species in 71 genera, it is the largest family of lichen-forming fungi. The most speciose genera in the family are the well-known groups: '' Xanthoparmelia'' ( 82 .... References nadvornikiana Lichen species Lichens described in 1932 Lichens of Europe Lichens of North America Taxa named by Vilmos Kőfaragó-Gyelnik {{Parmeliaceae-stub ...
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Clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population, or a species (extinct or Extant taxon, extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed ''monophyletic'' (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming Taxon, taxa that are not clades; that is, taxa that are not Monophyly, monophyletic. Some of the relationships between organisms that the molecul ...
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Gyrophoric Acid
Gyrophoric acid is a tridepside. It is a double ester of the orsellinic acid. It can also be found in most of the species of the lichen genera '' Actinogyra'', '' Lasallia'', and '' Umbilicaria'' . Natural occurrence and biosynthesis Gyrophoric acid is an orcinol-derived tridepside, meaning it is built from three orsellinic acid units linked by ester bonds. In nature the compound reaches its highest concentrations in rock-dwelling '' Umbilicaria'' lichens, but smaller amounts occur in several other genera, including '' Cryptothecia'', ''Xanthoparmelia'', '' Actinogyra'' and '' Lasallia''. Gyrophoric acid often co-exists with simpler didepsides such as lecanoric acid, suggesting a shared metabolic origin. Pharmacologically, gyrophoric acid has been demonstrated to possess broad-spectrum antimicrobial activity and a marked cytotoxic effect against a range of cancer cell lines in in vitro laboratory tests; it also absorbs ultraviolet radiation and has historically served as a ...
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Norstictic Acid
Norstictic acid is a depsidone produced as a secondary metabolites in lichens. The compound contains both an aldehyde carbonyl group and an adjacent hydroxyl In chemistry, a hydroxy or hydroxyl group is a functional group with the chemical formula and composed of one oxygen atom covalently bonded to one hydrogen atom. In organic chemistry, alcohols and carboxylic acids contain one or more hydroxy ... group in its molecular structure, which enables it to form complexes with certain metals. This property was demonstrated in studies of lichens growing on copper-rich substrates, where norstictic acid forms complexes with copper ions in the (outer layer) of the lichen thallus, resulting in a distinctive green-yellow coloration. The formation of these copper-norstictic acid complexes appears to be selective, as evidenced by comparing it to the structurally similar stictic acid, which lacks the adjacent hydroxy group and does not form copper complexes. This metal-binding capabi ...
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Variety (botany)
In botanical nomenclature, variety (abbreviated var.; in ) is a taxonomic rank below that of species and subspecies, but above that of form. As such, it gets a three-part infraspecific name. It is sometimes recommended that the subspecies rank should be used to recognize geographic distinctiveness, whereas the variety rank is appropriate if the taxon is seen throughout the geographic range of the species. Example The pincushion cactus, ''Escobaria vivipara'', is a wide-ranging variable species occurring from Canada to Mexico, and found throughout New Mexico below about . Nine varieties have been described. Where the varieties of the pincushion cactus meet, they intergrade. The variety ''Escobaria vivipara'' var. ''arizonica'' is from Arizona, while ''Escobaria vivipara'' var. ''neo-mexicana'' is from New Mexico. Definitions The term is defined in different ways by different authors. However, the International Code of Nomenclature for Cultivated Plants, while recognizing ...
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Cascade Mountains
The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as many of those in the North Cascades, and the notable volcanoes known as the High Cascades. The small part of the range in British Columbia is referred to as the Canadian Cascades or, locally, as the Cascade Mountains. The highest peak in the range is Mount Rainier in Washington at . part of the Pacific Ocean's Ring of Fire, the ring of volcanoes and associated mountains around the Pacific Ocean. All of the eruptions in the contiguous United States over the last 200 years have been from the Cascade Volcanoes. The two most recent were Lassen Peak from 1914 to 1921 and a major eruption of Mount St. Helens in 1980. Minor eruptions of Mount St. Helens have also occurred since, most recently from 2004 to 2008. The Cascade Range is a part of the American Cordil ...
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Washington (state)
Washington, officially the State of Washington, is a U.S. state, state in the Pacific Northwest region of the United States. It is often referred to as Washington State to distinguish it from Washington, D.C., the national capital, both named after George Washington (the first President of the United States, U.S. president). Washington borders the Pacific Ocean to the west, Oregon to the south, Idaho to the east, and shares Canada–United States border, an international border with the Provinces and territories of Canada, Canadian province of British Columbia to the north. Olympia, Washington, Olympia is the List of capitals in the United States, state capital, and the most populous city is Seattle. Washington is the List of U.S. states and territories by area, 18th-largest state, with an area of , and the List of U.S. states and territories by population, 13th-most populous state, with a population of just less than 8 million. The majority of Washington's residents live ...
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Oregon
Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of its eastern boundary with Idaho. The 42nd parallel north, 42° north parallel delineates the southern boundary with California and Nevada. The western boundary is formed by the Pacific Ocean. Oregon has been home to many Indigenous peoples of the Americas, indigenous nations for thousands of years. The first European traders, explorers, and settlers began exploring what is now Oregon's Pacific coast in the early to mid-16th century. As early as 1564, the Spanish expeditions to the Pacific Northwest, Spanish began sending vessels northeast from the Philippines, riding the Kuroshio Current in a sweeping circular route across the northern part of the Pacific. In 1592, Juan de Fuca undertook detailed mapping a ...
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Bruce McCune
Bruce Pettit McCune (born 1952) is an American lichenologist, botanist, plant ecologist, and software developer for analysis of ecological data. Biography McCune grew up in Cincinnati. He completed his freshman year of college at Lawrence University in Appleton, Wisconsin, and then transferred to the University of Montana in the autumn of 1971. There he graduated in 1974 with a bachelor's degree in botany. From 1971 to 1974 McCune and his then girlfriend, Patricia S. Muir, spent considerable time on Mount Sentinel, where they investigated lichens, mosses, and other plants. From 1974 to 1975 he travelled and also worked for two summers in Montana for the Bureau of Land Management. From 1976 to 1979 he was a graduate student at the University of Montana, where he graduated with a master's degree. In August 1979 he married Patricia Muir. She graduated in 1975 with a bachelor's degree in botany from the University of Montana. In 1979 the couple matriculated as graduate students at the ...
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Secondary Metabolite
Secondary metabolites, also called ''specialised metabolites'', ''secondary products'', or ''natural products'', are organic compounds produced by any lifeform, e.g. bacteria, archaea, fungi, animals, or plants, which are not directly involved in the normal cell growth, growth, Biological development, development, or reproduction of the organism. Instead, they generally mediate ecological biological interaction, interactions, which may produce a Natural selection, selective advantage for the organism by increasing its survivability or fecundity. Specific secondary metabolites are often restricted to a narrow set of species within a phylogenetic group. Secondary metabolites often play an important role in plant defense against herbivory and other interspecies defenses. Humans use secondary metabolites as medicines, flavourings, pigments, and recreational drugs. The term secondary metabolite was first coined by Albrecht Kossel, the 1910 Nobel Prize laureate for medicine and physio ...
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Genetic Variation
Genetic variation is the difference in DNA among individuals or the differences between populations among the same species. The multiple sources of genetic variation include mutation and genetic recombination. Mutations are the ultimate sources of genetic variation, but other mechanisms, such as genetic drift, contribute to it, as well. Among individuals within a population Genetic variation can be identified at many levels. Identifying genetic variation is possible from observations of phenotypic variation in either quantitative traits (traits that vary continuously and are coded for by many genes, e.g., leg length in dogs) or discrete traits (traits that fall into discrete categories and are coded for by one or a few genes, e.g., white, pink, or red petal color in certain flowers). Genetic variation can also be identified by examining variation at the level of enzymes using the process of protein electrophoresis. Polymorphic genes have more than one allele at each locu ...
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