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Apheloria Tigana
''Apheloria tigana'', Yellow-and-black Flat Millipede, is a large North American flat-backed millipede in the family Xystodesmidae. It is reported to secrete cyanide compounds as a defense. It is recommended that one wash hands after handling this organism as the toxic compounds it secretes are poisonous and can cause extreme irritation if rubbed in the eyes. Characteristics include yellow paranota (lateral segmental expansions on the dorsa), a yellow mid-dorsal spot on the anterior margin of the collum or 1st segment, and yellow mid-dorsal spots on the caudal-most 3-5 segments. South of the Cape Fear River basin there is an undescribed ''Apheloria'' species with yellow middorsal marks on most segments. ''A. tigana'' occurs in the Eastern United States, from southeastern North Carolina northward throughout the Blue Ridge Mountains.Hoffman, Richard L. 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publications 8, p. 294 R ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Myriapoda
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. The fossil record of myriapods reaches back into the late Silurian, although molecular evidence suggests a diversification in the Cambrian Period, and Cambrian fossils exist which resemble myriapods. The oldest unequivocal myriapod fossil is of the millipede '' Pneumodesmus newmani'', from the late Silurian (428 million years ago). ''P. newmani'' is also important as the earliest known terrestrial animal. The phylogenetic classification of myriapods is still debated. The scientific study of myriapods is myriapodology, and those who study myriapods are myriapodologists. Anatomy Myriapods have a single pair of antennae and, in most cases, simple eyes. Exceptions are the two classes symphylans and pauropods, and the millipede order Polydesmida and the centipede order Geophilomorpha, which are all ...
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Millipede
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of '' Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fu ...
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Polydesmida
Polydesmida (from the Greek ''poly'' "many" and ''desmos'' "bond") is the largest order of millipedes, containing approximately 3,500 species, including all the millipedes reported to produce hydrogen cyanide (HCN). Description Members of the order Polydesmida are also known as "flat-backed millipedes", because on most species, each body segment has wide lateral keels known as paranota. These keels are produced by the posterior half (metazonite) of each body ring behind the collum. Polydesmids have no eyes, and vary in length from . Many of the larger species show bright coloration patterns which warn predators of their toxic secretions. Adults usually have 20 segments, counting the collum as the first ring and the telson as the last ring. Juveniles have from 7 to 19 rings. In species with the usual 20 segments, adult females have 31 pairs of legs, but in adult males, the eighth leg pair (the first leg pair of the 7th ring) is modified into a single pair of gonopods, leaving on ...
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Xystodesmidae
Xystodesmidae is a family of millipedes. Its members often have very small distributional areas, with many species only known from a single locality. They are found across the northern hemisphere, with peak diversity in the Appalachian Mountains, where one-third of the 300 or so species occur. They are particularly abundant in deciduous broadleaf forests in the Mediterranean Basin, Africa, Asia, Central and North America, and Russia. Information on basic taxonomy is scant for this family; for example, it is estimated that the genus ''Nannaria'' contains over 200 species, but only 25 were described as of 2006. By 2022, 78 species in ''Nannaria'' have been described. Xystodesmidae contains many colorful and distinctive species, including ''Apheloria virginiensis'' of the eastern U.S. and '' Harpaphe haydeniana'' of the western U.S. The Sierra luminous millipedes of the genus ''Motyxia'' exhibit the only known examples of bioluminescence in the Polydesmida. Species of ''Apheloria'' ...
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Apheloria
''Apheloria'' is a genus of flat-backed millipedes in the family Xystodesmidae, occurring in the central and southeastern United States, and ranging as far north as southern Quebec, Canada.Hoffman, Richard L. 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publications 8, pp. 293-296Marek, Paul; Tanabe, Tsutomu; Sierwald, Petra (2014''A Species Catalog of the Millipede Family Xystodesmidae (Diplopoda: Polydesmida)''.Special Publication 17. Virginia Museum of Natural History. pp. 7-9 They are aposematically colored in black and contrasting reds and yellows, and some species in the Appalachian Mountains resemble species of '' Brachoria'' where they co-occur, a phenomenon known as Müllerian mimicry Müllerian mimicry is a natural phenomenon in which two or more well-defended species, often foul-tasting and sharing common predators, have come to mimicry, mimic each other's honest signal, honest aposematism, warning signals, ...
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Ralph Vary Chamberlin
Ralph Vary Chamberlin (January 3, 1879October 31, 1967) was an American biologist, ethnographer, and historian from Salt Lake City, Utah. He was a faculty member of the University of Utah for over 25 years, where he helped establish the School of Medicine and served as its first dean, and later became head of the zoology department. He also taught at Brigham Young University and the University of Pennsylvania, and worked for over a decade at the Museum of Comparative Zoology at Harvard University, where he described species from around the world. Chamberlin was a prolific taxonomist who named over 4,000 new animal species in over 400 scientific publications. He specialized in arachnids (spiders, scorpions, and relatives) and myriapods (centipedes, millipedes, and relatives), ranking among the most prolific arachnologists and myriapodologists in history. He described over 1,400 species of spiders, 1,000 species of millipedes, and the majority of North American centipedes, ...
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Flat-backed Millipede
Polydesmida (from the Greek ''poly'' "many" and ''desmos'' "bond") is the largest order of millipedes, containing approximately 3,500 species, including all the millipedes reported to produce hydrogen cyanide (HCN). Description Members of the order Polydesmida are also known as "flat-backed millipedes", because on most species, each body segment has wide lateral keels known as paranota. These keels are produced by the posterior half (metazonite) of each body ring behind the collum. Polydesmids have no eyes, and vary in length from . Many of the larger species show bright coloration patterns which warn predators of their toxic secretions. Adults usually have 20 segments, counting the collum as the first ring and the telson as the last ring. Juveniles have from 7 to 19 rings. In species with the usual 20 segments, adult females have 31 pairs of legs, but in adult males, the eighth leg pair (the first leg pair of the 7th ring) is modified into a single pair of gonopods, leaving ...
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Cyanide
Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of a carbon atom triple-bonded to a nitrogen atom. In inorganic cyanides, the cyanide group is present as the anion . Soluble salts such as sodium cyanide (NaCN) and potassium cyanide (KCN) are highly toxic. Hydrocyanic acid, also known as hydrogen cyanide, or HCN, is a highly volatile liquid that is produced on a large scale industrially. It is obtained by acidification of cyanide salts. Organic cyanides are usually called nitriles. In nitriles, the group is linked by a covalent bond to carbon. For example, in acetonitrile (), the cyanide group is bonded to methyl (). Although nitriles generally do not release cyanide ions, the cyanohydrins do and are thus rather toxic. Bonding The cyanide ion is isoelectronic with carbon m ...
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Blue Ridge Mountains
The Blue Ridge Mountains are a Physiographic regions of the world, physiographic province of the larger Appalachian Mountains range. The mountain range is located in the Eastern United States, and extends 550 miles southwest from southern Pennsylvania through Maryland, West Virginia, Virginia, North Carolina, South Carolina, Tennessee, and Georgia (U.S. state), Georgia. This province consists of northern and southern physiographic regions, which divide near the Roanoke River gap. To the west of the Blue Ridge, between it and the bulk of the Appalachians, lies the Great Appalachian Valley, bordered on the west by the Ridge-and-valley Appalachians, Ridge and Valley province of the Appalachian range. The Blue Ridge Mountains are known for having a bluish color when seen from a distance. Trees put the "blue" in Blue Ridge, from the isoprene released into the atmosphere. This contributes to the characteristic haze on the mountains and their perceived color. Within the Blue Ridge prov ...
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BugGuide
BugGuide (or BugGuide.net) is a website and online community of naturalists, both amateur and professional, who share observations of arthropods such as insects, spiders, and other related creatures. The website consists of informational guide pages and many thousands of photographs of arthropods from the United States and Canada which are used for identification and research. The non-commercial site is hosted by the Iowa State University Iowa State University of Science and Technology (Iowa State University, Iowa State, or ISU) is a public land-grant research university in Ames, Iowa. Founded in 1858 as the Iowa Agricultural College and Model Farm, Iowa State became one of the ... Department of Entomology. BugGuide was conceived by photographer Troy Bartlett in 2003 and since 2006 has been maintained by John VanDyk, an adjunct assistant professor of entomology and a senior systems analyst at Iowa State University.
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