Zoology (/zoʊˈɒlədʒi, zu-/)[note 1] is the branch of
biology that studies the animal kingdom, including the structure,
embryology, evolution, classification, habits, and distribution of all
animals, both living and extinct, and how they interact with their
ecosystems. The term is derived from
Ancient Greek ζῷον, zōion,
i.e. "animal" and λόγος, logos, i.e. "knowledge,
1.1 Ancient history to Darwin
3 Branches of zoology
4 See also
7 External links
Ancient history to Darwin
Conrad Gesner (1516–1565). His Historiae animalium is considered
the beginning of modern zoology.
Main article: History of zoology (through 1859)
The history of zoology traces the study of the animal kingdom from
ancient to modern times. Although the concept of zoology as a single
coherent field arose much later, the zoological sciences emerged from
natural history reaching back to the biological works of Aristotle and
Galen in the ancient Greco-Roman world. This ancient work was further
developed in the
Middle Ages by Muslim physicians and scholars such as
Albertus Magnus. During the
Renaissance and early modern period, zoological thought was
Europe by a renewed interest in empiricism and the
discovery of many novel organisms. Prominent in this movement were
Vesalius and William Harvey, who used experimentation and careful
observation in physiology, and naturalists such as Carl Linnaeus,
Jean-Baptiste Lamarck, and Buffon who began to classify the diversity
of life and the fossil record, as well as the development and behavior
Microscopy revealed the previously unknown world of
microorganisms, laying the groundwork for cell theory. The
growing importance of natural theology, partly a response to the rise
of mechanical philosophy, encouraged the growth of natural history
(although it entrenched the argument from design).
Over the 18th, 19th, and 20th centuries, zoology became an
increasingly professional scientific discipline. Explorer-naturalists
Alexander von Humboldt
Alexander von Humboldt investigated the interaction between
organisms and their environment, and the ways this relationship
depends on geography, laying the foundations for biogeography, ecology
and ethology. Naturalists began to reject essentialism and consider
the importance of extinction and the mutability of species. Cell
theory provided a new perspective on the fundamental basis of
Main article: History of zoology (since 1859)
These developments, as well as the results from embryology and
paleontology, were synthesized in Charles Darwin's theory of evolution
by natural selection. In 1859, Darwin placed the theory of organic
evolution on a new footing, by his discovery of a process by which
organic evolution can occur, and provided observational evidence that
it had done so.
Darwin gave a new direction to morphology and physiology, by uniting
them in a common biological theory: the theory of organic evolution.
The result was a reconstruction of the classification of animals upon
a genealogical basis, fresh investigation of the development of
animals, and early attempts to determine their genetic relationships.
The end of the 19th century saw the fall of spontaneous generation and
the rise of the germ theory of disease, though the mechanism of
inheritance remained a mystery. In the early 20th century, the
rediscovery of Mendel's work led to the rapid development of genetics,
and by the 1930s the combination of population genetics and natural
selection in the modern synthesis created evolutionary
Cell biology studies the structural and physiological properties of
cells, including their behavior, interactions, and environment. This
is done on both the microscopic and molecular levels, for
single-celled organisms such as bacteria as well as the specialized
cells in multicellular organisms such as humans. Understanding the
structure and function of cells is fundamental to all of the
biological sciences. The similarities and differences between cell
types are particularly relevant to molecular biology.
Anatomy considers the forms of macroscopic structures such as organs
and organ systems. It focuses on how organs and organ
systems work together in the bodies of humans and animals, in addition
to how they work independently.
Anatomy and cell biology are two
studies that are closely related, and can be categorized under
Animal anatomical engraving from Handbuch der Anatomie der Tiere
Physiology studies the mechanical, physical, and biochemical processes
of living organisms by attempting to understand how all of the
structures function as a whole. The theme of "structure to function"
is central to biology. Physiological studies have traditionally been
divided into plant physiology and animal physiology, but some
principles of physiology are universal, no matter what particular
organism is being studied. For example, what is learned about the
physiology of yeast cells can also apply to human cells. The field of
animal physiology extends the tools and methods of human physiology to
Physiology studies how for example nervous, immune,
endocrine, respiratory, and circulatory systems, function and
Evolutionary research is concerned with the origin and descent of
species, as well as their change over time, and includes scientists
from many taxonomically oriented disciplines. For example, it
generally involves scientists who have special training in particular
organisms such as mammalogy, ornithology, herpetology, or entomology,
but use those organisms as systems to answer general questions about
Evolutionary biology is partly based on paleontology, which uses the
fossil record to answer questions about the mode and tempo of
evolution, and partly on the developments in areas such as
population genetics and evolutionary theory. Following the
development of DNA fingerprinting techniques in the late 20th century,
the application of these techniques in zoology has increased the
understanding of animal populations. In the 1980s,
developmental biology re-entered evolutionary biology from its initial
exclusion from the modern synthesis through the study of evolutionary
developmental biology. Related fields often considered
part of evolutionary biology are phylogenetics, systematics, and
Scientific classification in zoology, is a method by which zoologists
group and categorize organisms by biological type, such as genus or
Biological classification is a form of scientific taxonomy.
Modern biological classification has its root in the work of Carl
Linnaeus, who grouped species according to shared physical
characteristics. These groupings have since been revised to improve
consistency with the Darwinian principle of common descent. Molecular
phylogenetics, which uses
DNA sequences as data, has driven many
recent revisions and is likely to continue to do so. Biological
classification belongs to the science of zoological systematics.
Linnaeus's table of the animal kingdom from the first edition of
Systema Naturae (1735).
Many scientists now consider the five-kingdom system outdated. Modern
alternative classification systems generally start with the
Archaea (originally Archaebacteria); Bacteria
(originally Eubacteria); Eukaryota (including protists, fungi, plants,
and animals) These domains reflect whether the cells have
nuclei or not, as well as differences in the chemical composition of
the cell exteriors.
Further, each kingdom is broken down recursively until each species is
separately classified. The order is:
Domain; kingdom; phylum; class; order; family; genus; species. The
scientific name of an organism is generated from its genus and
species. For example, humans are listed as Homo sapiens. Homo is the
genus, and sapiens the specific epithet, both of them combined make up
the species name. When writing the scientific name of an organism, it
is proper to capitalize the first letter in the genus and put all of
the specific epithet in lowercase. Additionally, the entire term may
be italicized or underlined.
The dominant classification system is called the Linnaean taxonomy. It
includes ranks and binomial nomenclature. The classification,
taxonomy, and nomenclature of zoological organisms is administered by
the International Code of Zoological Nomenclature. A merging draft,
BioCode, was published in 1997 in an attempt to standardize
nomenclature, but has yet to be formally adopted.
Kelp gull chicks peck at red spot on mother's beak to stimulate the
Ethology is the scientific and objective study of animal behavior
under natural conditions, as opposed to behaviourism,
which focuses on behavioral response studies in a laboratory setting.
Ethologists have been particularly concerned with the evolution of
behavior and the understanding of behavior in terms of the theory of
natural selection. In one sense, the first modern ethologist was
Charles Darwin, whose book, The Expression of the Emotions in Man and
Animals, influenced many future ethologists.
Biogeography studies the spatial distribution of organisms on the
Earth, focusing on topics like plate tectonics, climate
change, dispersal and migration, and cladistics. The creation of this
study is widely accredited to Alfred Russel Wallace, a British
biologist who had some of his work jointly published with Charles
Branches of zoology
Although the study of animal life is ancient, its scientific
incarnation is relatively modern. This mirrors the transition from
natural history to biology at the start of the 19th century. Since
Hunter and Cuvier, comparative anatomical study has been associated
with morphography, shaping the modern areas of zoological
investigation: anatomy, physiology, histology, embryology, teratology
and ethology. Modern zoology first arose in German and
British universities. In Britain,
Thomas Henry Huxley
Thomas Henry Huxley was a prominent
figure. His ideas were centered on the morphology of animals. Many
consider him the greatest comparative anatomist of the latter half of
the 19th century. Similar to Hunter, his courses were composed of
lectures and laboratory practical classes in contrast to the previous
format of lectures only.
Gradually zoology expanded beyond Huxley's comparative anatomy to
include the following sub-disciplines:
Zoography, also known as descriptive zoology, is the applied science
of describing animals and their habitats
Comparative anatomy studies the structure of animals
Ethology studies animal behavior
The various taxonomically oriented disciplines such as mammalogy,
biological anthropology, herpetology, ornithology, ichthyology, and
entomology identify and classify species and study the structures and
mechanisms specific to those groups.
Evolutionary biology: Development of both animals and plants is
considered in the articles on evolution, population genetics,
heredity, variation, Mendelism, and reproduction.
Molecular biology studies the common genetic and developmental
mechanisms of animals and plants
Systematics, cladistics, phylogenetics, phylogeography, biogeography,
and taxonomy classify and group species via common descent and
Animal science, the biology of domesticated animals
List of zoologists
Outline of zoology
Timeline of zoology
^ The pronunciation of zoology as /zuˈɒlədʒi/ is typically
regarded as nonstandard, though is not uncommon.
^ "zoology". Online Etymology Dictionary..mw-parser-output
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^ Lois N. Magner (2002). A History of the Life Sciences, Revised and
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^ Jan Sapp (2003). "Chapter 7". Genesis: The
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^ William Coleman (1978). "Chapter 2".
Biology in the Nineteenth
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^ Jerry A. Coyne (2009). Why
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^ Henry Gray (1918).
Anatomy of the
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^ John H. Gillespie (1998).
Population Genetics: A Concise Guide.
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^ Vassiliki Betty Smocovitis (1996). Unifying Biology: The
Evolutionary Synthesis and Evolutionary Biology. Princeton University
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^ Heather Silyn-Roberts (2000). Writing for Science and Engineering:
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^ John McNeill (4 November 1996). "The BioCode: Integrated biological
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At Wikiversity, you can learnmore and teach others about
the School of Zoology.
Wikibooks has more on the topic of: Zoology
Wikisource has original works on the topic: Zoology
Zoology at Project Gutenberg
Online Dictionary of Invertebrate Zoology
.mw-parser-output .nobold font-weight:normal Domain
Ctenophora (comb jellies)
Cnidaria (jellyfish and relatives)
Echinodermata (starfish and relatives)
Kinorhyncha (mud dragons)
Priapulida (penis worms)
Nematomorpha (horsehair worms)
Loricifera (corset animals)
Onychophora (velvet worms)
Chaetognatha (arrow worms)
Gnathostomulida (jaw worms)
Rotifera (wheel animals)
Acanthocephala (thorny-headed worms)
Dicyemida or Rhombozoa
Annelida (ringed worms)
Nemertea (ribbon worms)
Entoprocta or Kamptozoa
Ectoprocta (moss animals)
Brachiopoda (lamp shells)
Phoronida (horseshoe worms)
Major groupswithin phyla
Anthozoa inc. corals
Medusozoa inc. jellyfish
Asterozoa inc. starfish
Phyla with ≥1000 extant species bolded
Evolutionary developmental biology
Hierarchy of life
Ecosystem > Community (Biocoenosis)
Organism > Organ system
> Organ > Tissue > Cell > Organelle
Biomolecular complex >
Biomolecule) > Atom
Earliest known life forms
Plant morphology terms
Karl Ernst von Baer
Jakob von Uexküll
Alfred Russel Wallace
Timeline of zoology
vteZoos, aquariums and aviariesTypes of zoos
Animal theme park
Marine mammal park
Ex situ conservation
In situ conservation
Animals in captivity