''Yurgovuchia'' (meaning "coyote") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the ...

that lived in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

during the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145& ...

period in what is now the

Cedar Mountain Formation The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid- Cretaceous. The formation was named for Cedar Mountain in northern Emery County, Utah, where Will ...

. It contains a single species, ''Yurgovuchia doellingi''. The remains were discovered in

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...

Discovery and naming

The holotype was collected by Donald D. DeBlieux in 2005, from Don's Place, part of the Doelling's Bowl bone bed in Grand County, Utah. This bone bed is in the lower Yellow Cat Member of the

, dating probably to the

Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretace ...

stage of the

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

, about 139–134.6

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago ...

. ''Yurgovuchia'' was first described and named by Phil Senter,

James I. Kirkland James Ian Kirkland (born August 24, 1954) is an American Palaeontologist, paleontologist and geologist. He has worked with dinosaur remains from the south west United States of America and Mexico and has been responsible for discovering new and i ...

, Donald D. DeBlieux, Scott Madsen and Natalie Toth in

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gat ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

is ''Yurgovuchia doellingi''. The generic name is derived from the Ute word ''yurgovuch'', meaning

coyote The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological nich ...

, a predator of similar size to ''Y. doellingi'' which currently inhabits the same region. The specific name, ''doellingi'', honors the geologist Helmut Doelling, for his 50-plus years of geological research and mapping of Utah for the Utah Geological Survey and for causing the discovery of the Doelling's Bowl dinosaur sites, in which ''Y. doellingi'' was collected.

Description

''Yurgovuchia'' is known only from a single individual represented by an associated partial

postcranial Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

and the only known specimen, UMNH VP 20211, includes cervical,

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...

, and

caudal vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

as well as the proximal end of a left pubis. The preserved vertebrae are fused, indicating that the specimen was an adult at the time of death and not a juvenile specimen of ''

Utahraptor ''Utahraptor'' (meaning "Utah's thief") is a genus of large dromaeosaurid dinosaur that lived in North America during the Early Cretaceous period. It was a heavy-built, ground-dwelling, bipedal carnivore. It contains a single species, ''Utahrap ...

''. Instead of this, it represents a smaller and different species of

, with an estimed size of .

According to its describers in 2012, ''Yurgovuchia'' can be recognized from other dromaeosaurid taxa by the following characteristics: each side of the

axial Axial may refer to: * one of the anatomical directions describing relationships in an animal body * In geometry: :* a geometric term of location :* an axis of rotation * In chemistry, referring to an axial bond * a type of modal frame, in music * ...

centrum have a single pneumatopore, third cervical vertebra with cranial end of the centrum not beveled, cervical

prezygapophyses The articular processes or zygapophyses (Greek ζυγον = "yoke" (because it links two vertebrae) + απο = "away" + φυσις = "process") of a vertebra are projections of the vertebra that serve the purpose of fitting with an adjacent vertebr ...

is flexed, cervical vertebrae epipophyses is above the postzygapophyseal facets, the pubis lacks pubic

tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...

, the cranial faces of the centrum of caudal vertebrae are round, the cervical and dorsal vertebrae preserve hypapophyses without pneumatopores, the caudal prezygapophyses is distally elongated to the transition point, but not surpassing the length of a centrum.

Classification

According to the

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...

analysis performed by its describers in 2012, ''Yurgovuchia'' represents advanced

dromaeosaurine Dromaeosaurinae is a subfamily of the theropod group Dromaeosauridae. The earliest dromaeosaurine is ''Utahraptor'', dating back to the Early Cretaceous period in North America, however, some isolated teeth seems to represent an indeterminate spe ...

s, closely related to '' Achillobator'', '' Dromaeosaurus'' and ''

''. Below are the results obtained:

Paleoecology

Cedar Mountain Formation Yellow Cat Fauna

Other dinosaurs are also known from Don's Place including the iguanodontians ''

Iguanacolossus ''Iguanacolossus'' (meaning "Iguana Colossus" or "Colossal Iguana") is a genus of iguanodontian ornithopod dinosaur that lived in North America during the Early Cretaceous period. It is known from UMNH VP 20205, the associated holotype with a lar ...

'',

polacanthine Polacanthinae is a subfamily of ankylosaurs, most often nodosaurids, from the Late Jurassic through Early Cretaceous of Europe and potentially North America and Asia. The group is defined as the largest clade closer to ''Polacanthus foxii'' than ...

s, and

velociraptorine Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably ''Nuthetes'' from the United Kingdom, and possibly '' Deinonychus'' from North America. However, several indeterminate velociraptori ...

dromaeosaurids represented by a pubis (UMNH VP 21752) and a tentative radius (UMNH VP 21751). Many additional theropods have previously been described from the Yellow Cat Member, including the therizinosaur '' Falcarius'' and the small, predatory troodontids '' Geminiraptor'' from the lower part of the member, in addition, the large dromaeosaurines ''

'', small

ornithomimosaurs Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and Nor ...

'' Nedcolbertia'', and unnamed

eudromaeosaurs Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were relatively small to medium-sized, feathered hypercarnivores (with diets consisting almost entirely of other terrestrial vertebrates) ...

represented by a tail skeleton (UMNH VP 20209) from the upper part of the member. ''Yurgovuchia'' shared its environment and lived alongside other dinosaurs in the Lower Yellow Cat, such as the

theropods Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...

''Falcarius'' and ''Geminiraptor'', the

sauropods Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...

Mierasaurus ''Mierasaurus'' is an extinct genus of sauropod dinosaur from the Early Cretaceous of Utah, United States. The taxon was first described and named in 2017 by Rafael Royo-Torres and colleagues, from a mostly complete skeleton including a disarti ...

'' and the large iguanodontians ''Iguanacolossus''. There are also indeterminate

goniopholidid Goniopholididae is an extinct family of moderate-sized semi-aquatic neosuchian crocodyliformes. Their bodyplan and morphology are convergent on living crocodilians. They lived across Laurasia (Asia, Europe and North America) between the Middle Ju ...

crocodiles and the unnamed velociraptorines known from the Lower Yellow Cat.

References

{{Taxonbar, from=Q3236057 Eudromaeosaurs Early Cretaceous dinosaurs of North America Fossil taxa described in 2012 Taxa named by James I. Kirkland Paleontology in Utah

Discovery and naming

Description

Classification

Paleoecology

See also

References