Walking catfish
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The walking catfish (''Clarias batrachus'') is a species of freshwater airbreathing catfish native to
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
. It is named for its ability to "walk" and wiggle across dry land, to find food or suitable environments. While it does not truly walk as most bipeds or quadrupeds do, it has the ability to use its
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
s to keep it upright as it makes a wiggling motion with snakelike movements to traverse land. This fish normally lives in slow-moving and often stagnant waters in ponds, swamps, streams, and rivers, as well as in flooded rice paddies, or temporary pools that may dry up. When this happens, its "walking" skill allows the fish to move to other aquatic environments. Considerable taxonomic confusion surrounds this species and it has frequently been confused with other close relatives.Ng, Heok Hee, and Kottelat, Maurice (2008). ''The identity of Clarias batrachus (Linnaeus, 1758), with the designation of a neotype (Teleostei: Clariidae).'' Zoological Journal of the Linnean Society 153: 725–732. One main distinction between the walking catfish and the native North American ictalurid catfish with which it sometimes is confused, is that the walking catfish lacks an adipose fin.


Characteristics and anatomy

The walking catfish has an elongated body shape and reaches almost in length and in weight. Often covered laterally in small white spots, the body is mainly coloured a gray or grayish brown. This catfish has long-based dorsal and
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
s, as well as several pairs of sensory barbels. The skin is scaleless, but covered with
mucus Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It ...
, which protects the fish when it is out of water. This fish needs to be handled carefully when fishing it due to its embedded sting or thorn-like defensive mechanism hidden behind its fins (including the middle ones before the tail fin, similar to the majority of all catfishes).


Taxonomy, distribution, and habitat

The walking catfish is a
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
species native to
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
. The native range of true ''Clarias batrachus'' is confirmed from the
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Gui ...
n island of
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
only, but three closely related and more widespread species frequently have been confused with this species. These are '' C. magur'' of northeast India and
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...
, a likely undescribed species from
Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...
, and another likely undescribed species from the
Thai-Malay Peninsula The Malay Peninsula ( Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The ar ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, and
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and e ...
. Both of these undescribed species have been referred to as, ''Clarias ''
aff. ''Species affinis'' (commonly abbreviated to: ''sp.'', ''aff.'', or ''affin.'') is taxonomic terminology in zoology and botany. In open nomenclature it indicates that available material or evidence suggests that the proposed species is related to, h ...
'' batrachus''. At present, the taxonomic position of the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
population (called ''hito'' or simply "catfish" by the locals) is unclear, and it also is unclear whether South Indian populations are ''C. magur'' or another species. As a consequence, much information (behavioral, ecological, related to introduced populations, etc.) listed for ''C. batrachus'', may be for the closely related species that have been confused with true ''C. batrachus''. True ''C. batrachus'', ''C. magur'' and the two likely undescribed species are all kept in
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
. Walking catfish thrive in stagnant, frequently hypoxic waters, and often are found in muddy
pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from tha ...
s,
canal Canals or artificial waterways are waterways or engineered channels built for drainage management (e.g. flood control and irrigation) or for conveyancing water transport vehicles (e.g. water taxi). They carry free, calm surface f ...
s,
ditch A ditch is a small to moderate divot created to channel water. A ditch can be used for drainage, to drain water from low-lying areas, alongside roadways or fields, or to channel water from a more distant source for plant irrigation. Ditches ar ...
es, and similar habitats. The species spends most of its time on, or right above, the bottom, with occasional trips to the surface to gulp air.


Diet

In the wild, this creature is
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
; it feeds on smaller fish,
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is est ...
s, and other
invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordat ...
, as well as
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts comm ...
and aquatic weeds. It is a voracious eater that consumes food supplies rapidly, so it is considered harmful when invasive.


As an invasive species

Within Asia, this species has been introduced widely. It has also been introduced outside its native range where it is considered an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...
. It consumes the food supplies of native fish and preys on their young. It also is regarded as an invasive species because they can destroy fish farms. In the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
, it is established in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
. It is reported in
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
,
Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
,
Massachusetts Massachusetts (Massachusett: ''Muhsachuweesut Massachusett_writing_systems.html" ;"title="nowiki/> məhswatʃəwiːsət.html" ;"title="Massachusett writing systems">məhswatʃəwiːsət">Massachusett writing systems">məhswatʃəwiːsət'' En ...
, and
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
. The walking catfish was imported to Florida, reportedly from
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
, in the early 1960s for the
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
trade. The first introductions apparently occurred in the mid-1960s when adult fish imported as brood stock escaped, either from a fish farm in northeastern Broward County or from a truck transporting brood fish between Dade and Broward Counties. Additional introductions in Florida, supposedly purposeful releases, were made by fish farmers in the
Tampa Bay Tampa Bay is a large natural harbor and shallow estuary connected to the Gulf of Mexico on the west-central coast of Florida, comprising Hillsborough Bay, McKay Bay, Old Tampa Bay, Middle Tampa Bay, and Lower Tampa Bay. The largest freshwater ...
area, Hillsborough County in late 1967 or early 1968, after the state banned the importation and possession of walking catfish. Aquarium releases likely are responsible for introductions in other states. Dill and Cordone (1997) reported this species has been sold by tropical fish dealers in California for some time. They also have been spotted occasionally in the Midwest. In Florida, walking catfish are known to have invaded aquaculture farms, entering ponds where they prey on agricultural fish stocks. In response, fish farmers have had to erect fences to protect ponds. Authorities have also created laws that ban possession of walking catfish. In 2017, ''Clarias'' spp. were recovered from the
River Tonge The River Tonge is a short river, splitting Bolton from contiguous Tonge, both in Greater Manchester, England. The Tonge is formed at the ''Meeting of the Waters'', where Astley Brook, from Smithills in the west, meets the Eagley Brook drawi ...
, near
Bolton Bolton (, locally ) is a large town in Greater Manchester in North West England, formerly a part of Lancashire. A former mill town, Bolton has been a production centre for textiles since Flemish weavers settled in the area in the 14th ...
, Northern England.


As food

In Thailand, this fish is known as, ''pla duk dan'' ( th, ปลาดุกด้าน). It is a common, inexpensive food item, prepared in a variety of ways. It often is offered by street vendors, especially grilled or fried. This fish is one of the most common freshwater catfish in the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
where it is known as ''hito'' in the local language. In Indonesia, it is called ''lele'', and it is the main ingredient in several native dishes, such as ''
pecel lele Pecel lele or pecak lele is an Indonesian deep-fried ''Clarias'' catfish dish originating from Lamongan, East Java, Indonesia. Dish It consists of catfish served with traditional ''sambal'' chili paste, often served with fried tempeh and/or tof ...
''. Not true ''C. batrachus'', but ''C. magur'' is eaten in
West Bengal West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fou ...
as magur curry ( bn, মাগুর মাছের ঝোল), and is considered good during illness, particularly for body weakness. It is prepared in a light curry sauce with coriander powder and cinnamon powder. It reportedly is fed to children to develop body strength. In
Karnataka Karnataka (; ISO 15919, ISO: , , also known as Karunāḍu) is a States and union territories of India, state in the southwestern region of India. It was Unification of Karnataka, formed on 1 November 1956, with the passage of the States Reor ...
, ''C. magur'' is called ''murgodu'' (ಮುರ‍್ಗೋಡು). In coastal Karnataka, it is called ''mugudu'' ( tcy, ಮುಗುಡು), and is considered a delicacy. Neither the Thai nor the Indian populations are likely to be ''C. batrachus'', however.


Aquarium

A white variation with black patterns is commonly seen in the aquarium fish trade. However, this color variation also is prohibited where walking catfish are banned. Very well-rooted plants and large structures that provide some shade should be included in an aquarium with these fish. Any small tankmates will be eaten by this fish.


See also

* Eel catfish *
List of freshwater aquarium fish species A vast number of aquatic species have successfully adapted to live in the freshwater aquarium. This list gives some examples of the most common species found in home aquariums. Catfish Characins and other characiformes ...
*
List of invasive species in the Everglades Invasive species in the Everglades are exotic plants and animals that are not native to the area and have aggressively adapted to conditions in wilderness areas in southern Florida. The Everglades are a massive watershed in the southern portio ...


References


External links


Ros, Wolfgang
Datz 57 (7): 12–15. * [http://www.catfish-and-more.com/ Ros, Wolfgang

Datz 59 (4): 33–37. {{DEFAULTSORT:Walking Catfish Clarias Fish of Bangladesh Fish of Southeast Asia Fish of Thailand Fish described in 1758 Taxa named by Carl Linnaeus Fish of the Philippines Amphibious fish pam:Itu