Typha
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''Typha'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of about 30
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
monocotyledon Monocotyledons (), commonly referred to as monocots, ( Lilianae '' sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one ...
ous
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s in the family Typhaceae. These plants have a variety of common names, in
British English British English (BrE, en-GB, or BE) is, according to Oxford Dictionaries, "English as used in Great Britain, as distinct from that used elsewhere". More narrowly, it can refer specifically to the English language in England, or, more broadl ...
as bulrush or reedmace, in
American English American English, sometimes called United States English or U.S. English, is the set of varieties of the English language native to the United States. English is the most widely spoken language in the United States and in most circumstances ...
as reed, cattail, or punks, in
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
as cumbungi or bulrush, in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
as bulrush or cattail, and in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
as raupo. Other taxa of plants may be known as bulrush, including some sedges in '' Scirpus'' and related genera. The genus is largely distributed in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, where it is found in a variety of
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (Anoxic waters, anoxic) processes prevailing, especially in t ...
habitats. The
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
s are edible. Evidence of preserved
starch Starch or amylum is a polymeric carbohydrate consisting of numerous glucose units joined by glycosidic bonds. This polysaccharide is produced by most green plants for energy storage. Worldwide, it is the most common carbohydrate in human die ...
grains on grinding stones suggests they were already eaten in Europe 30,000 years ago.


Description

''Typha'' are aquatic or semi-aquatic, rhizomatous, herbaceous
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s. The leaves are glabrous (hairless), linear, alternate and mostly basal on a simple, jointless stem that bears the flowering spikes. The plants are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is ...
, with unisexual flowers that develop in dense
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s. The numerous male flowers form a narrow spike at the top of the vertical stem. Each male (staminate) flower is reduced to a pair of
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s and hairs, and withers once the
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
is shed. Large numbers of tiny female flowers form a dense,
sausage A sausage is a type of meat product usually made from ground meat—often pork, beef, or poultry—along with salt, spices and other flavourings. Other ingredients, such as grains or breadcrumbs may be included as fillers or extenders. ...
-shaped spike on the stem below the male spike. In larger species this can be up to long and thick. The seeds are minute, long, and attached to fine hairs. When ripe, the heads disintegrate into a cottony fluff from which the seeds disperse by wind.


General ecology

''Typha'' are often among the first wetland plants to colonize areas of newly exposed wet mud, with their abundant wind-dispersed seeds. Buried seeds can survive in the soil for long periods of time. They germinate best with sunlight and fluctuating temperatures, which is typical of many wetland plants that regenerate on mud flats. The plants also spread by
rhizomes In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
, forming large, interconnected stands. ''Typha'' are considered to be dominant competitors in wetlands in many areas, and they often exclude other plants with their dense canopy. In the bays of the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
, for example, they are among the most abundant wetland plants. Different species of cattails are adapted to different water depths. Well-developed aerenchyma make the plants tolerant of submersion. Even the dead stalks are capable of transmitting oxygen to the rooting zone. Although ''Typha'' are native wetland plants, they can be aggressive in their competition with other native species. They have been problematic in many regions in North America, from the Great Lakes to the Everglades. Native sedges are displaced and wet meadows shrink, likely as a response to altered hydrology of the wetlands and increased nutrient levels. An introduced or hybrid species may be contributing to the problem. Control is difficult. The most successful strategy appears to be mowing or burning to remove the aerenchymous stalks, followed by prolonged flooding. It may be more important to prevent invasion by preserving water level fluctuations, including periods of drought, and to maintain infertile conditions. ''Typha'' are frequently eaten by wetland mammals such as
muskrat The muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over a wide range of climates and habita ...
s, which also use them to construct feeding platforms and dens, thereby also providing nesting and resting places for waterfowl.


Accepted species and natural hybrids

The following names are currently accepted: The most widespread species is ''Typha latifolia'', which is distributed across the entire temperate northern hemisphere. It has also been introduced to Australia. ''T. angustifolia'' is nearly as widespread, but does not extend as far north; it may be introduced and invasive in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
. ''T. domingensis'' has a more southern
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
distribution, and it occurs in Australia. ''T. orientalis'' is widespread in Asia, Australia, and New Zealand. ''T. laxmannii'', ''T. minima'', and ''T. shuttleworthii'' are largely restricted to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
and southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.


Uses


Culinary

Many parts of the ''Typha'' plant are edible to humans. Before the plant flowers, the tender inside of the shoots can be squeezed out and eaten raw or cooked. The starchy
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
s are nutritious with a protein content comparable to that of maize or rice. They can be processed into a
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many cul ...
with 266 kcal per 100 grams, and are most often harvested from late autumn to early spring. They are fibrous, and the starch must be scraped or sucked from the tough fibers. Baby shoots emerging from the rhizomes, which are sometimes subterranean, can be picked and eaten raw. Also underground is a
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...
lump which can be peeled and eaten raw or cooked like a
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Uni ...
. The plant is one championed by survival experts because various parts can be eaten throughout the year. Plants growing in polluted water can accumulate
lead Lead is a chemical element with the symbol Pb (from the Latin ) and atomic number 82. It is a heavy metal that is denser than most common materials. Lead is soft and malleable, and also has a relatively low melting point. When freshly cut, ...
and
pesticide Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and ...
residues in their rhizomes, and these should not be eaten. The rind of young stems can be peeled off, and the tender white heart inside can be eaten raw or boiled and eaten like asparagus. This food has been popular among the
Cossacks The Cossacks , es, cosaco , et, Kasakad, cazacii , fi, Kasakat, cazacii , french: cosaques , hu, kozákok, cazacii , it, cosacchi , orv, коза́ки, pl, Kozacy , pt, cossacos , ro, cazaci , russian: казаки́ or ...
in Russia, and has been called "Cossack asparagus". The leaf bases can be eaten raw or cooked, especially in late spring when they are young and tender. In early summer the sheath can be removed from the developing green flower spike, which can then be boiled and eaten like
corn on the cob Corn on the cob is a culinary term for a cooked ear of sweet corn (maize) eaten directly off the cob. The ear is picked while the endosperm is in the "milk stage" so that the kernels are still tender. Ears of corn are steamed, boiled, or ...
. In mid-summer when the male flowers are mature, the
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
can be collected and used as a flour supplement or thickener.


Agriculture

The seeds have a high
linoleic acid Linoleic acid (LA) is an organic compound with the formula COOH(CH2)7CH=CHCH2CH=CH(CH2)4CH3. Both alkene groups are ''cis''. It is a fatty acid sometimes denoted 18:2 (n-6) or 18:2 ''cis''-9,12. A linoleate is a salt or ester of this acid. ...
content and can be used to feed cattle and chickens. They can also be found in African countries like Ghana. Harvesting cattail removes nutrients from the wetland that would otherwise return via the decomposition of decaying plant matter. Floating mats of cattails remove nutrients from eutrophied bodies of freshwater.


Building material

For local native tribes around Lake Titicaca in
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
and
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
, ''Typha'' were among the most important plants and every part of the plant had multiple uses. For example, they were used to construct rafts and other boats. During World War II, the United States Navy used the down of ''Typha'' as a substitute for kapok in life vests and aviation jackets. Tests showed that even after 100 hours of submersion, the buoyancy was still effective. ''Typha'' are used as
thermal insulation Thermal insulation is the reduction of heat transfer (i.e., the transfer of thermal energy between objects of differing temperature) between objects in thermal contact or in range of radiative influence. Thermal insulation can be achieved with ...
in buildings as an organic alternative to conventional insulating materials such as
glass wool Glass wool is an insulating material made from glass fiber arranged using a binder into a texture similar to wool. The process traps many small pockets of air between the glass, and these small air pockets result in high thermal insulation pro ...
or stone wool.


Paper

''Typha'' stems and leaves can be used to make
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distribu ...
. It is strong with a heavy texture and it is hard to bleach, so it is not suitable for industrial production of graphical paper. In 1853, considerable amounts of cattail paper were produced in
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
, due to a shortage of raw materials. In 1948, French scientists tested methods for annual harvesting of the leaves. Because of the high cost, these methods were abandoned and no further research was done. Today ''Typha'' is used to make decorative paper.


Fiber

Fibers up to 4 meters long can be obtained from the stems when they are treated mechanically or chemically with
sodium hydroxide Sodium hydroxide, also known as lye and caustic soda, is an inorganic compound with the formula NaOH. It is a white solid ionic compound consisting of sodium cations and hydroxide anions . Sodium hydroxide is a highly caustic base and al ...
. The stem fibers resemble
jute Jute is a long, soft, shiny bast fiber that can be spun into coarse, strong threads. It is produced from flowering plants in the genus ''Corchorus'', which is in the mallow family Malvaceae. The primary source of the fiber is '' Corchorus ol ...
and can be used to produce raw textiles. The leaf fibers can be used as an alternative to
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor pe ...
and
linen Linen () is a textile made from the fibers of the flax plant. Linen is very strong, absorbent, and dries faster than cotton. Because of these properties, linen is comfortable to wear in hot weather and is valued for use in garments. It also ...
in clothing. The yield of leaf fiber is 30 to 40 percent and ''Typha glauca'' can produce 7 to 10 tons per hectare annually.


Biofuel

''Typha'' can be used as a source of starch to produce
ethanol Ethanol (abbr. EtOH; also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound. It is an alcohol with the chemical formula . Its formula can be also written as or (an ethyl group linked to a ...
. Because of their high productivity in northern latitudes, ''Typha'' are considered to be a bioenergy crop.


Other

The seed hairs were used by some
indigenous peoples of the Americas The Indigenous peoples of the Americas are the inhabitants of the Americas before the arrival of the European settlers in the 15th century, and the ethnic groups who now identify themselves with those peoples. Many Indigenous peoples of the A ...
as
tinder Tinder is easily combustible material used to start a fire. Tinder is a finely divided, open material which will begin to glow under a shower of sparks. Air is gently wafted over the glowing tinder until it bursts into flame. The flaming tinder i ...
for starting fires. Some tribes also used ''Typha'' down to line
moccasins A moccasin is a shoe, made of deerskin or other soft leather, consisting of a sole (made with leather that has not been "worked") and sides made of one piece of leather, stitched together at the top, and sometimes with a vamp (additional panel o ...
, and for bedding, diapers, baby powder, and
cradleboard Cradleboards (, se, gietkka, sms, ǩiõtkâm, smn, kietkâm, sje, gietkam) are traditional protective baby-carriers used by many indigenous cultures in North America and throughout northern Scandinavia amongst the Sámi. There are a variety ...
s. One Native American word for ''Typha'' meant "fruit for papoose's bed". ''Typha'' down is still used in some areas to stuff clothing items and pillows. ''Typha'' can be dipped in wax or fat and then lit as a candle, the stem serving as a wick. Without the use of wax or fat it will smolder slowly, somewhat like
incense Incense is aromatic biotic material that releases fragrant smoke when burnt. The term is used for either the material or the aroma. Incense is used for aesthetic reasons, religious worship, aromatherapy, meditation, and ceremony. It may also b ...
, and may repel insects. The flower stalks can be made into
chopsticks Chopsticks ( or ; Pinyin: ''kuaizi'' or ''zhu'') are shaped pairs of equal-length sticks of Chinese origin that have been used as kitchen and eating utensils in most of East and Southeast Asia for over three millennia. They are held in the ...
. The leaves can be treated to weave into baskets, mats, or sandals. The rushes are harvested and the leaves often dried for later use in chair seats. Re-wetted, the leaves are twisted and wrapped around the chair rungs to form a densely woven seat that is then stuffed (usually with the left over rush). Small-scale experiments have indicated that ''Typha'' are able to remove
arsenic Arsenic is a chemical element with the symbol As and atomic number 33. Arsenic occurs in many minerals, usually in combination with sulfur and metals, but also as a pure elemental crystal. Arsenic is a metalloid. It has various allotropes, b ...
from drinking water. The boiled rootstocks have been used as a
diuretic A diuretic () is any substance that promotes diuresis, the increased production of urine. This includes forced diuresis. A diuretic tablet is sometimes colloquially called a water tablet. There are several categories of diuretics. All diuretics i ...
for increasing urination, or mashed to make a jelly-like paste for sores, boils, wounds, burns, scabs, and smallpox pustules. Cattail pollen is used as a banker source of food for predatory insects and mites (such as ''
Amblyseius swirskii ''Typhlodromips swirskii'', the Swirski mite (synonymy: ''Amblyseius swirskii'', ''Amblyseius rykei'', ''Neoseiulus swirskii''), is a species of predatory mite in the family Phytoseiidae. It is used in biological pest control of western flower t ...
'') in greenhouses. The cattail, or, as it is commonly referred to in the American Midwest, the sausage tail, has been the subject of multiple artist renditions, gaining popularity in the mid-twentieth century. The term, sausage tail, derives from the similarity that cattails have with sausages, a name given to the plant by the Midwest Polish community who had noticed a striking similarity between the plant and a common Polish dish, kiełbasa.


See also

*
Typha domingensis ''Typha domingensis'', known commonly as southern cattail or cumbungi, is a perennial herbaceous plant of the genus ''Typha''. Distribution and habitat It is found throughout temperate and tropical regions worldwide. It is sometimes found as a s ...


References


External links


Can you actually eat cattails?
from '' The Straight Dope'' {{Authority control Aquatic plants Inflorescence vegetables Poales genera Root vegetables