Tritylodon
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''Tritylodon'' (Greek for 3 cusped tooth) is an extinct genus of
tritylodonts Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, bearing several mammalian traits like erect limbs, endothermy and details o ...
, one of the most advanced group of cynodont therapsids. They lived in the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
and possibly Late Triassic periods along with
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s. They also shared many characteristics with mammals, and were once considered mammals because of overall skeleton construction. That was changed due to them retaining the vestigial reptilian jawbones and a different skull structure. Tritylodons are now regarded as non-mammalian
synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptil ...
s.


Characteristics

If a living ''Tritylodon'' were to be seen today, it would look a lot like a large rodent. They were about long but there is no certainty about the exact weight. Their method of chewing food, a grinding motion with the bottom teeth sliding against the top teeth, resembled that of rodents as well. The bottom teeth were much like a set of cusps and the top teeth were a set of matching grooves that matched perfectly allowing this motion. There were large incisors at the very front of their mouth separated by a gap from the rest of the teeth. The incisors would stick out and remain slightly visible when the mouth was closed. The legs were directly beneath the body like mammals, unlike the earlier therapsids with sprawling limbs. These animals were burrowers; the structure of the shoulder, front limbs, and large front incisors show this. They used their incisors to help dig and unearth buried plants. The way they ate and the shape of their teeth demonstrate that ''Tritylodons'' were probably primarily
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
(though some tritylodontids show evidence of more omnivorous diets, and modern analogues like rodents tend to be more omnivorous than their dentitions lead on). Any of the Tritylodonts including ''Tritylodon'' were
warm-blooded Warm-blooded is an informal term referring to animal species which can maintain a body temperature higher than their environment. In particular, homeothermic species maintain a stable body temperature by regulating metabolic processes. The onl ...
or endothermic. Like most non-
placental Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguishe ...
mammalimorphs, it had
epipubic Epipubic bones are a pair of bones projecting forward from the pelvic bones of modern marsupials, monotremes and fossil mammals like multituberculates, and even basal eutherians (the ancestors of placental mammals, who lack them). They first o ...
bones, aiding in its erect gait but preventing the expansion of the abdomen, making it unable to go through prolonged pregnancy and instead give birth to larval young like modern marsupials and
monotremes Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals (Eutheria), and marsupials ( Metatheria). Monotremes are typified by structural differences in their br ...
.


Habitat

The ''Tritylodons habitat was limited to the forests of South Africa, with other fossils found in the
Hanson Formation The Hanson Formation (also known as the Shafer Peak Formation) is a geologic formation on Mount Kirkpatrick and north Victoria Land, Antarctica. It is one of the two major dinosaur-bearing rock groups found on Antarctica to date; the other is the ...
of Antarctica. When the species originated, about 200 million years ago, the African area was drier and hotter. But for most of their existence the climate was tropical and wetter.


Fossils

The ''Tritylodon'' fossils in South Africa are found concentrated mainly in an area about 11,000 km² (4,250 mi²). They have been found in floodplain deposits of the Lower Jurassic
Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, south ...
(upper
Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a peri ...
). In this area there have been so many findings it has been named the Tritylodon Acme Zone. The fossil findings have all been in the Free State of South Africa. The genus ''Tritylodon'' of the Tritylodonts is restricted to the South African forms: ''Tritylodon longaevus'' and ''Tritylodon maximus''. It is suggested that ''T. maximus'' is either a large ''T. longeavus'' or a closely related species. If it is a closely related species it could possibly be ecological succession since the larger ''T. maximus'' fossils have been dated in the Sinemurian–Pliensbachian mainly less than 190 million years ago and the ''T. longaevus'' in the Hettangian–Sinemurian mainly more than 190 million years ago. With the fossil findings of each species overlapping in Sinemurian stage, the fossils show two differences, ''T. maximus'' being larger and having nine upper postcanines (neither species had canine teeth) instead of the seven teeth like ''T. longeavus''. All other structures of the two ''Tritylodon'' species were the same.http://home.arcor.de/ktdykes/jtherap.htm#tritylodon Below is a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships:


References


External links

* * * * *https://archive.today/20130204222846/http://tritylodontidae.totallyexplained.com/ * {{Taxonbar, from=Q1441437 Prehistoric cynodont genera Jurassic synapsids of Africa Fossil taxa described in 1884 Taxa named by Richard Owen Tritylodontids