Tree squirrel
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Tree squirrels are the members of the
squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
(Sciuridae) commonly just referred to as "squirrels." They include more than 100 arboreal species native to all continents except
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
and
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
. They do not form a single natural, or monophyletic, group; they are variously related to others in the squirrel family, including ground squirrels, flying squirrels, marmots, and chipmunks. The defining characteristic used to determine which species of Sciuridae are tree squirrels is dependent on their
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
rather than their
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemic ...
. Tree squirrels live mostly among trees, as opposed to those that live in
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of s ...
s in the ground or among rocks. An exception is the flying squirrel that also makes its home in trees, but has a physiological distinction separating it from its tree squirrel cousins: special flaps of skin called patagia, acting as glider wings, which allow gliding flight. The best known
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of tree squirrels is ''
Sciurus The genus ''Sciurus'' contains most of the common, bushy-tailed squirrels in North America, Europe, temperate Asia, Central America and South America. Species The number of species in the genus is subject to change. In 2005, Thorington & H ...
'', which includes the eastern gray squirrel of North America ( introduced to
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It ...
in the 1870s), the
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbe ...
of Eurasia, and the North American fox squirrel, among many others. Many tree squirrel species have adapted to human-altered environments such as rural farms, suburban backyards and urban parks.


Classification

Current taxonomy, based on genetic data, splits the tree squirrels into several subfamilies. The following genera of the squirrel family are classified as tree squirrels. *Subfamily Ratufinae **Genus '' Ratufa'' (Asian giant squirrels) *Subfamily
Sciurillinae The Neotropical pygmy squirrel (''Sciurillus pusillus'') is a South American species of tree squirrel, being the only living species in the genus ''Sciurillus'' and the subfamily Sciurillinae. Genetic analysis has shown it to be the sister group ...
**Genus '' Sciurillus'' (South American pygmy squirrel) *Subfamily
Sciurinae Sciurinae is a subfamily of squirrels (in the family Sciuridae), uniting the flying squirrels with certain related tree squirrels. Older sources place the flying squirrels in a separate subfamily (Pteromyinae) and unite all remaining sciurids into ...
**Tribe Sciurini (mostly American tree squirrels) ***Genus ''
Microsciurus ''Microsciurus'' or dwarf squirrels is a genus of squirrels from the tropical regions of Central and South America. Taxonomy and systematics Recent DNA analysis has shown that there is some confusion regarding the traditional classification ...
'' (American dwarf squirrels) ***Genus '' Rheithrosciurus'' (Borneo tufted ground squirrel) ***Genus ''
Sciurus The genus ''Sciurus'' contains most of the common, bushy-tailed squirrels in North America, Europe, temperate Asia, Central America and South America. Species The number of species in the genus is subject to change. In 2005, Thorington & H ...
'' (Eurasian and American tree squirrels) ***Genus ''
Syntheosciurus Bangs's mountain squirrel (''Syntheosciurus brochus'') is a poorly known species of tree squirrel, that only lives in Costa Rica and Panama. It can be found in mountain rain forests at an altitude between , and lives mainly in the tree tops, bu ...
'' (Central American mountain squirrel) ***Genus ''
Tamiasciurus Pine squirrels are squirrels of the genus ''Tamiasciurus'', in the Sciurini tribe, of the large family Sciuridae. Species This genus includes three species: *''Tamiasciurus hudsonicus'' — American red squirrel *''Tamiasciurus douglasii'' ...
'' (American pine squirrels) *Subfamily Callosciurinae (Asian tree squirrels) **Genus ''
Callosciurus ''Callosciurus'' is a genus of squirrels collectively referred to as the "''beautiful squirrels''". They are found mainly in Southeast Asia, though a few species also occur in Nepal, northeastern India, Bangladesh and southern China. Several of t ...
'' (Oriental tree squirrels, introduced into Europe and South America) **Genus ''
Exilisciurus ''Exilisciurus'' is a genus of rodent in the family Sciuridae Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chi ...
'' (Asian pygmy squirrels) **Genus '' Funambulus'' (Asian palm squirrels, introduced into Australia in the 1920s) **Genus '' Glyphotes'' (sculptor squirrel) **Genus '' Nannosciurus'' (Asian dwarf squirrel) **Genus ''
Prosciurillus ''Prosciurillus'' is a genus of rodent in the family Sciuridae, endemic to Sulawesi and nearby small islands, Indonesia. It contains the following species: * Secretive dwarf squirrel (''Prosciurillus abstrusus'') * Alston’s Sulawesi dwarf sq ...
'' (Sulawesi dwarf squirrels) **Genus '' Rubrisciurus'' (Sulawesi giant squirrel) **Genus ''
Sundasciurus ''Sundasciurus'' is a genus of rodent in the family Sciuridae Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chi ...
'' (Sunda squirrels) **Genus '' Tamiops'' (Asian striped squirrels) *Subfamily Xerinae **Tribe Protoxerini (African tree squirrels) ***Genus '' Epixerus'' (African palm squirrels) ***Genus '' Funisciurus'' (rope squirrels) ***Genus '' Heliosciurus'' (sun squirrels) ***Genus '' Myosciurus'' (African pygmy squirrel) ***Genus '' Paraxerus'' (bush squirrels) ***Genus '' Protoxerus'' (African giant squirrels)


Relationship with humans

Squirrels are generally inquisitive and persistent animals. In residential neighborhoods, they are notorious for tenaciously trying to circumvent obstacles in order to eat from bird feeders. Although they are expert climbers, and primarily arboreal, some species of squirrels also thrive in urban environments, where they have adapted to humans.


As pests

Squirrels are sometimes considered
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
because of their propensity to chew on various edible and inedible objects, and their stubborn persistence in trying to get what they want. Their characteristic gnawing trait also aids in maintaining sharp
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, ...
, and because their teeth grow continuously, prevents their over-growth. On occasion, squirrels will chew through plastic and even metal to get to food. Tree squirrels may bury food in the ground for later retrieval. Squirrels use their keen sense of smell to search for buried food, but can dig numerous holes in the process. This may become an annoyance to
gardeners A gardener is someone who practices gardening, either professionally or as a hobby. Description A gardener is any person involved in gardening, arguably the oldest occupation, from the hobbyist in a residential garden, the home-owner suppleme ...
with strict landscape requirements, especially when the garden contains edibles. Homeowners in areas with a heavy squirrel population must be vigilant in keeping attics,
basements A basement or cellar is one or more Storey, floors of a building that are completely or partly below the storey, ground floor. It generally is used as a utility space for a building, where such items as the Furnace (house heating), furnace, ...
, and sheds carefully sealed to prevent property damage caused by nesting squirrels. A squirrel nest is called a " drey". Squirrels are a serious fire hazard when they break into buildings. They often treat exposed power cables as tree branches, and gnaw on the electrical insulation. The resulting exposed conductors can short out, causing a fire. For this reason alone, squirrel nests inside buildings cannot be safely ignored. A squirrel nest will also cause problems with noise, excreta, unpleasant odors, and eventual structural damage. Some homeowners resort to more interesting ways of dealing with this problem, such as collecting and placing fur from pets such as domestic
cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
and dogs in attics. It is hoped that this fur would indicate to nesting squirrels that a potential predator roams, and will encourage evacuation. Odoriferous repellents, including
mothballs Mothballs are small balls of chemical pesticide and deodorant, sometimes used when storing clothing and other materials susceptible to damage from mold or moth larvae (especially clothes moths like ''Tineola bisselliella''). Composition Ol ...
and
ammonia Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogenous ...
, are generally ineffective in expelling squirrels from buildings. Once established in a nest, squirrels stubbornly ignore fake owls and
scarecrow A scarecrow is a decoy or mannequin, often in the shape of a human. Humanoid scarecrows are usually dressed in old clothes and placed in open fields to discourage birds from disturbing and feeding on recently cast seed and growing crops.Lesle ...
s, along with bright flashing lights, loud noises, and ultrasonic or electromagnetic devices. However, squirrels must leave the nest to obtain food and water (usually daily, except in bad weather), affording an opportunity to trap them or exclude them from re-entering. To discourage chewing on an object, it can be coated or covered with something to make it distasteful: for instance a soft cloth doused with
chili pepper Chili peppers (also chile, chile pepper, chilli pepper, or chilli), from Nahuatl '' chīlli'' (), are varieties of the berry-fruit of plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for ...
paste or powder.
Capsaicin Capsaicin (8-methyl-''N''-vanillyl-6-nonenamide) ( or ) is an active component of chili peppers, which are plants belonging to the genus ''Capsicum''. It is a chemical irritant for mammals, including humans, and produces a sensation of burnin ...
and Ro-pel are other forms of repellent. To remain effective, the coating must be reapplied regularly, especially if it is exposed to the weather. Poisoning squirrels can be problematic because of the risks to other animals or children in the building, and because the odor of a dead squirrel in an attic or wall cavity is very unpleasant and persistent. Trapping is often necessary to remove squirrels from residential structures. Effective baits include fruit,
peanut butter Peanut butter is a food paste or spread made from ground, dry-roasted peanuts. It commonly contains additional ingredients that modify the taste or texture, such as salt, sweeteners, or emulsifiers. Peanut butter is consumed in many countri ...
, nuts, seeds and vanilla extract. An alternative method is to wait until squirrels have left in search of food, and then close up all their access openings, or to install one-way
trap doors A trapdoor is a sliding or hinged door in a floor or ceiling. It is traditionally small in size. It was invented to facilitate the hoisting of grain up through mills, however, its list of uses has grown over time. The trapdoor has played a pivot ...
or a carefully angled pipe. Attempting to get rid of all squirrels in a neighborhood is generally a futile goal; the focus instead should be on physically excluding them from places where they can do damage. There are other humane techniques to remove squirrels from buildings, but removal is ineffective unless steps are taken to prevent them from immediately breaking in again. Squirrels are often the cause of power outages. They can readily climb a power pole and crawl or run along a power cable. The animals will climb onto power
transformer A transformer is a passive component that transfers electrical energy from one electrical circuit to another circuit, or multiple circuits. A varying current in any coil of the transformer produces a varying magnetic flux in the transformer' ...
s or
capacitor A capacitor is a device that stores electrical energy in an electric field by virtue of accumulating electric charges on two close surfaces insulated from each other. It is a passive electronic component with two terminals. The effect of ...
s looking for food, or a place to cache acorns. If they touch a
high voltage High voltage electricity refers to electrical potential large enough to cause injury or damage. In certain industries, ''high voltage'' refers to voltage above a certain threshold. Equipment and conductors that carry high voltage warrant sp ...
conductor and a
ground Ground may refer to: Geology * Land, the surface of the Earth not covered by water * Soil, a mixture of clay, sand and organic matter present on the surface of the Earth Electricity * Ground (electricity), the reference point in an electrical c ...
ed portion of the enclosure at the same time, they are
electrocuted Electrocution is death or severe injury caused by electric shock from electric current passing through the body. The word is derived from "electro" and "execution", but it is also used for accidental death. The term "electrocution" was coined ...
, and often cause a
short circuit A short circuit (sometimes abbreviated to short or s/c) is an electrical circuit that allows a current to travel along an unintended path with no or very low electrical impedance. This results in an excessive current flowing through the circu ...
that shuts down equipment. Squirrels have brought down the high-tech
NASDAQ The Nasdaq Stock Market () (National Association of Securities Dealers Automated Quotations Stock Market) is an American stock exchange based in New York City. It is the most active stock trading venue in the US by volume, and ranked second ...
stock market twice and were responsible for a spate of power outages at the
University of Alabama The University of Alabama (informally known as Alabama, UA, or Bama) is a public research university in Tuscaloosa, Alabama. Established in 1820 and opened to students in 1831, the University of Alabama is the oldest and largest of the publ ...
. To sharpen their teeth, squirrels will often chew on tree branches or even the occasional live power line. Rubber or plastic plates, or freely rotating sleeves ("squirrel guards") are sometimes used to discourage access to these facilities. Squirrels otherwise appear to be safe and pose almost zero risk of transmitting
rabies Rabies is a viral disease that causes encephalitis in humans and other mammals. Early symptoms can include fever and tingling at the site of exposure. These symptoms are followed by one or more of the following symptoms: nausea, vomiting, ...
. Squirrels cause economic losses to homeowners, nut growers, and forest managers in addition to damage to electric transmission lines. These losses include direct damage to property, repairs, lost revenue and public relations. While dollar costs of these losses are sometimes calculated for isolated incidents, there is no tracking system to determine the total extent of the losses.


As roadkill and traffic hazards

In regions where squirrels are plentiful, tire-flattened roadkill is a common sight on roadways, especially in the spring and fall, when there is a fresh crop of young rodents. Motorists have caused serious accidents by attempting to swerve or stop to avoid a squirrel in the road. Evasive maneuvers are difficult since squirrels are much more agile and have much quicker reaction times than motorists in heavy vehicles; the majority of vehicular encounters end with no harm to either party. An effort to mitigate these hazards to both squirrels and humans is the Nutty Narrows Bridge in Longview, Washington, listed on the
National Register of Historic Places The National Register of Historic Places (NRHP) is the United States federal government's official list of districts, sites, buildings, structures and objects deemed worthy of preservation for their historical significance or "great artistic ...
. It provides a way for squirrels to cross a busy street safely.


As urban wildlife

Tree squirrels are a common type of
urban wildlife Urban wildlife is wildlife that can live or thrive in urban/suburban environments or around densely populated human settlements such as townships. Some urban wildlife, such as house mice, are synanthropic, ecologically associated with and ev ...
. They can be trained to be hand-fed and will take as much food as is available because they cache the surplus. Squirrels living in parks and
campus A campus is traditionally the land on which a college or university and related institutional buildings are situated. Usually a college campus includes libraries, lecture halls, residence halls, student centers or dining halls, and park-li ...
es in cities have learned that humans are typically a ready source of food. Urban squirrels have learned to get a lot of food from generous or unknowingly 'careless' humans. Humans commonly offer various nuts and seeds; however, wildlife rehabilitators in the field have noted that neither raw nor roasted
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics, important to both small an ...
s nor
sunflower seed The sunflower seed is the seed of the sunflower (''Helianthus annuus''). There are three types of commonly used sunflower seeds: linoleic (most common), high oleic, and sunflower oil seeds. Each variety has its own unique levels of monounsat ...
s are healthy for squirrels, because they are deficient in several essential nutrients. This type of deficiency has been found to cause metabolic bone disease, a somewhat common ailment found in malnourished squirrels.


As food


In the US

In many areas of the U.S. squirrels are still hunted for food, as they were historically.It tasted like chicken
Retrieved December 19, 2008.
Recipes calling for squirrel even appear in cookbooks, including ''James Beard's American Cookery'' and pre-1997 copies of '' The Joy of Cooking''. Squirrel meat can be substituted for
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit sp ...
or
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adu ...
in many recipes and was an ingredient in the original recipe for
Brunswick stew Brunswick stew is a tomato-based stew generally involving local beans, vegetables, and originally small game meat such as squirrel or rabbit, though today often chicken. The exact origin of the stew is disputed. The states of Virginia, North Caro ...
, a popular dish in various parts of the Southern U.S. Other similar stews were also based on squirrel meat, including burgoo and
Southern Illinois chowder Southern Illinois Chowder is a thick stew or soup, very different from the New England and Manhattan chowders. In Edwards County, Illinois, it refers to both the food and to the social gathering at which it is prepared and served. It is belie ...
. Although squirrel meat is low in fat content, unlike most game meat it has been found by the
American Heart Association The American Heart Association (AHA) is a nonprofit organization in the United States that funds cardiovascular medical research, educates consumers on healthy living and fosters appropriate cardiac care in an effort to reduce disability and deat ...
to be high in
cholesterol Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol (or modified steroid), a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell memb ...
. Squirrels Unlimited host a World Championship Squirrel cook-off each year in Bentonville, Arkansas.


In the UK

For most of the history of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and ...
, squirrel has been a meat not commonly eaten, and even scorned by many. In the early 21st century however, wild squirrel has become a more popular meat to cook with, showing up in restaurants and shops more often in Britain as a fashionable alternative meat. Specifically, Britons are cooking with the invasive gray squirrel, which is praised for its low fat content and the fact that it comes from
free range Free range denotes a method of farming husbandry where the animals, for at least part of the day, can roam freely outdoors, rather than being confined in an enclosure for 24 hours each day. On many farms, the outdoors ranging area is fenced, ...
sources. Additionally, the novelty of a meat considered unusual or special has contributed to the spread of squirrel consumption. Due to the difficulty of a clean kill and other factors, the majority of squirrel eaten in the UK is acquired from professional hunters, trappers and gamekeepers. Some Britons are eating gray squirrel as a direct attempt to help the native
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbe ...
, which has been dwindling since the 19th-century introduction of the gray squirrel, resulting in dramatic
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
for the indigenous red squirrels. This factor was marketed by a national "Save Our Squirrels" campaign that used the slogan, "Save a red, eat a grey!"


Risks of eating

As with other wild game and
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
species, the consumption of squirrels that have been exposed to high levels of
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, th ...
or toxic waste poses a health risk to humans. In 2007 in the northern New Jersey community of Ringwood, the New Jersey Department of Health and Senior Services issued a warning to anyone who eats squirrel (especially children and those who are pregnant) to limit their consumption after a
lead Lead is a chemical element with the symbol Pb (from the Latin ) and atomic number 82. It is a heavy metal that is denser than most common materials. Lead is soft and malleable, and also has a relatively low melting point. When freshly cut, ...
-contaminated squirrel was found near the Ringwood Mines Landfill. Toxic waste had been illegally dumped at this location for many years, before authorities cracked down on this practice in the 1980s. In 1997, doctors in
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia ...
published a paper in the '' Lancet'' that considered a possible association between the local tradition of consuming squirrel brains and five cases of Creutzfeldt–Jakob disease, a rare but serious
prion Prions are misfolded proteins that have the ability to transmit their misfolded shape onto normal variants of the same protein. They characterize several fatal and transmissible neurodegenerative diseases in humans and many other animals. It i ...
-based disorder. The authors posed this as a mere possibility, unconfirmed by either post-mortem analysis of the patients' brain tissue, or identification of a contagious prion agent in squirrels. Nonetheless, the ''Lancet'' article generated substantial media coverage, including articles in the New Yorker and
New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
. A 2015 case of CJD in a
Pittsburgh Pittsburgh ( ) is a city in the Commonwealth (U.S. state), Commonwealth of Pennsylvania, United States, and the county seat of Allegheny County, Pennsylvania, Allegheny County. It is the most populous city in both Allegheny County and Wester ...
man who had eaten squirrel brains played out similarly: the media seized on the patient's unconventional food choice, positing squirrel brains as the source of his disease. The doctor who made the initial report later clarified that he had not meant to assert the squirrel meat was the cause. Analysis of the patient's brain tissue ruled out the possibility of CJD acquired from food. As of 2018, Creutzfeldt–Jakob disease had never been identified in squirrels, and the association between squirrel consumption and CJD remained speculative.


Relationship with trees

The biggest source of food for tree squirrels is tree nuts. Red squirrels store nuts in a single stash (a
midden A midden (also kitchen midden or shell heap) is an old dump for domestic waste which may consist of animal bone, human excrement, botanical material, mollusc shells, potsherds, lithics (especially debitage), and other artifacts and eco ...
) that tends to dry out, so the seeds don't take root. Fox squirrels and gray squirrels bury nuts over a widespread area ( scatterhoarding), and often forget them, resulting in new trees ( mutualism).


In culture

In the ''
Ramayana The ''Rāmāyana'' (; sa, रामायणम्, ) is a Sanskrit epic composed over a period of nearly a millennium, with scholars' estimates for the earliest stage of the text ranging from the 8th to 4th centuries BCE, and later stages ...
'', an ancient
Sanskrit Sanskrit (; attributively , ; nominally , , ) is a classical language belonging to the Indo-Aryan languages, Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had Trans-cultural diffusion ...
epic poem, a squirrel assists in constructing a bridge from India to
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
to help
Rama Rama (; ), Ram, Raman or Ramar, also known as Ramachandra (; , ), is a major deity in Hinduism. He is the seventh and one of the most popular '' avatars'' of Vishnu. In Rama-centric traditions of Hinduism, he is considered the Supreme Bei ...
rescue his wife
Sita Sita (; ) also called as Janaki and Vaidehi is a Hindu goddess and the female protagonist of the Hindu epic, ''Ramayana''. She is the consort of Rama, the avatar of the god Vishnu, and is regarded as a form of Vishnu's consort, Lakshmi. She ...
. Rama rewards the squirrel by stroking his back with his three middle fingers, thus giving the Indian palm squirrel the three white stripes that appear on its back. In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern per ...
, the squirrel
Ratatoskr In Norse mythology, Ratatoskr (Old Norse, generally considered to mean "drill-tooth"Orchard (1997:129), Simek (2007:261), and Byock (2005:173). or "bore-tooth"Lindow (2001:259).) is a squirrel who runs up and down the world tree Yggdrasil to carry ...
is a messenger who scurries up and down the trunk of the world-tree Yggdrasil, carrying malicious gossip and insults back and forth between the dragon
Níðhöggr In Norse mythology, Níðhöggr (''Malice Striker'', in Old Norse traditionally also spelled Níðhǫggr , often anglicized NidhoggWhile the suffix of the name, ''-höggr'', clearly means "striker" the prefix is not as clear. In particular, the ...
, who sits at the bottom of the tree gnawing on its roots, and the hawk Veðrfölnir, who sits at the top of the tree keeping watch. According to Richard W. Thorington, Jr. and Katie E. Ferrell, this legend may have originated from the red squirrel's habit of giving a "scolding alarm call in response to danger", which some Norsemen may have imagined as insults. In
Irish mythology Irish mythology is the body of myths native to the island of Ireland. It was originally oral tradition, passed down orally in the Prehistoric Ireland, prehistoric era, being part of ancient Celtic religion. Many myths were later Early Irish ...
, the goddess Medb is said to always have a bird perched on one shoulder and a squirrel on the other, serving as her messengers to the sky and the earth respectively. In Europe during the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
, squirrels were sometimes used in bestiaries as symbols of greed and avarice on account of their storing of nuts, but, in the nineteenth century, British natural history books often praised them as thrifty for this same reason. A myth told by the
Ainu people The Ainu are the indigenous people of the lands surrounding the Sea of Okhotsk, including Hokkaido Island, Northeast Honshu Island, Sakhalin Island, the Kuril Islands, the Kamchatka Peninsula and Khabarovsk Krai, before the arrival of the Ya ...
of
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
holds that squirrels are the discarded sandals of the ancestral deity Aioina, possibly because squirrels move in spurts like footsteps. The ''
Kalevala The ''Kalevala'' ( fi, Kalevala, ) is a 19th-century work of epic poetry compiled by Elias Lönnrot from Karelian and Finnish oral folklore and mythology, telling an epic story about the Creation of the Earth, describing the controversies and ...
'', a Finnish epic poem collected in the nineteenth century but rooted in much older
oral tradition Oral tradition, or oral lore, is a form of human communication wherein knowledge, art, ideas and Culture, cultural material is received, preserved, and transmitted orally from one generation to another.Jan Vansina, Vansina, Jan: ''Oral Traditio ...
, contains references to squirrels, including mention of a white squirrel being born of a virgin.
Literary Literature is any collection of written work, but it is also used more narrowly for writings specifically considered to be an art form, especially prose fiction, drama, and poetry. In recent centuries, the definition has expanded to includ ...
references to squirrels include the works of Beatrix Potter,
Brian Jacques James Brian Jacques (, as in "Jakes"; 15 June 1939 – 5 February 2011) was an English novelist known for his ''Redwall'' series of novels and ''Castaways of the Flying Dutchman'' series. He also completed two collections of short stories entit ...
' Redwall series (including Jess Squirrel and numerous other squirrels), Pattertwig in
C. S. Lewis Clive Staples Lewis (29 November 1898 – 22 November 1963) was a British writer and Anglican lay theologian. He held academic positions in English literature at both Oxford University (Magdalen College, 1925–1954) and Cambridge Univer ...
' '' Prince Caspian'', Michael Tod's Woodstock Saga of novels featuring squirrel communities in the style of ''
Watership Down ''Watership Down'' is an adventure novel by English author Richard Adams, published by Rex Collings Ltd of London in 1972. Set in Berkshire in southern England, the story features a small group of rabbits. Although they live in their natural ...
'', and the Starwife and her subjects from Robin Jarvis's Deptford novels. The title character in Miriam Young's 1964 children's book ''Miss Suzy'' is a squirrel.
Anthropomorphic Anthropomorphism is the attribution of human traits, emotions, or intentions to non-human entities. It is considered to be an innate tendency of human psychology. Personification is the related attribution of human form and characteristics t ...
red squirrels were used in British road safety campaigns between the 1950s and 1980s. An episode of the radio program ''
This American Life ''This American Life'' (''TAL'') is an American monthly hour-long radio program produced in collaboration with Chicago Public Media and hosted by Ira Glass. It is broadcast on numerous public radio stations in the United States and internati ...
'' called "Squirrel Cop" describes the unintentionally humorous misadventure of a newly hired policeman in trying to remove a frantic squirrel from a homeowner's living room, which results in personal injury and a small fire. First aired in 1998, this episode turned out to be one of the most popular ones of the series, prompting rebroadcasts and a lead position on the two-CD compilation '' Crimebusters + Crossed Wires: Stories from This American Life''.


Albino and white squirrels

One of the ways that squirrels affect human society is inspired by the fascination that people seem to have over local populations of white squirrels (often misidentified as being albino). This manifests itself by the creation of
social group In the social sciences, a social group can be defined as two or more people who interact with one another, share similar characteristics, and collectively have a sense of unity. Regardless, social groups come in a myriad of sizes and varieties ...
communities that form from a commonly shared interest in these rare animals. Other impacts on human society inspired by white squirrels include the creation of organizations that seek to protect them from human predation, and the use of the white squirrel image as a cultural icon. Although these squirrels are commonly referred to as "albinos", most of them are likely non-albino squirrels that exhibit a rare white fur coloration known as leucism that is as a result of a recessive gene found within certain eastern gray squirrel (''Sciurus carolinensis'') populations, and so technically they ought to be referred to as ''white'' squirrels, instead of ''albino''."Debunking the Albino Squirrel: Lack of white, pink-eyed critters on campus may disprove superstition" by Hudson Lockett, The Daily Texan, May 1, 2009. A project run by Untamed Science is seeking to report and document the occurrence of both white squirrels, albinos, and other piebald morphs. Users are encouraged to submit their sightings.The White and Albino Squirrel Phenomenon + New US Maps!
/ref> UNTalbinosquirrel.jpg, A true albino squirrel. Note the pink eyes. Albino squirrel Minneapolis 2018.jpg, Albino squirrel head on, showing blue irises Wsquirrel.png, A white squirrel. Note the non-pink eyes.


Local pride

Olney, Illinois Olney ( ) is the county seat in Richland County, Illinois. The population was 9,115 at the time of the 2010 census. History Settlement of the Richland County area began around 1815 when Thaddeus Morehouse, a native of Vermont, arrived by wa ...
, known as the "White Squirrel Capital of the World", is home of the world's largest known white squirrel colony. These squirrels have the right of way on all streets in the town, with a $500 fine for hitting one. The Olney Police Department features the image of a white squirrel on its officers' uniform patches. Along with Olney, there are four other towns in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
that avidly compete with each other to be the official "Home of the White Squirrel", namely:
Marionville, Missouri Marionville is a city in Lawrence County, Missouri, United States. The population was 2,225 at the 2010 census. Marionville is known for its large population of white squirrels. History Marionville was platted in 1854. The city name has been ...
;
Brevard, North Carolina Brevard is a city in Transylvania County, North Carolina, United States, with a population of 7,609 as of the 2010 Census. It is the county seat of Transylvania County. Brevard is located at the entrance to Pisgah National Forest and has becom ...
; Exeter, Ontario; and
Kenton, Tennessee Kenton is a town in Gibson and Obion counties, Tennessee. The population was 1,281 at the 2010 census, a decline of 25 from 2000. The Gibson County portion of Kenton is part of the Humboldt, TN Micropolitan Statistical Area, while the Obion Co ...
, each of which holds an annual white squirrel festival, among other things designed to promote their claim of "White Squirrel Capital". A list of white squirrel sightings around the world is maintained by the White Squirrel Research Institute, a group based in Brevard, North Carolina. Other towns that have reported white squirrel populations in North America (although not necessarily competing to be the "official" white squirrel capital) include Bowling Green, Kentucky; Columbia, Mississippi; DeForest, Wisconsin;
Stratford, Connecticut Stratford is a town in Fairfield County, Connecticut, United States. It is situated on Long Island Sound at the mouth of the Housatonic River. Stratford is in the Bridgeport–Stamford–Norwalk Metropolitan Statistical Area. It was settled ...
; and some of the snowbelt cities in the Western,
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known a ...
and Finger Lakes regions of New York State ( Buffalo, Rochester,
Ithaca Ithaca most commonly refers to: *Homer's Ithaca, an island featured in Homer's ''Odyssey'' *Ithaca (island), an island in Greece, possibly Homer's Ithaca *Ithaca, New York, a city, and home of Cornell University and Ithaca College Ithaca, Ithaka ...
and
Syracuse Syracuse may refer to: Places Italy * Syracuse, Sicily, or spelled as ''Siracusa'' * Province of Syracuse United States *Syracuse, New York **East Syracuse, New York ** North Syracuse, New York * Syracuse, Indiana *Syracuse, Kansas *Syracuse, M ...
). The Trinity Bellwoods neighborhood of Toronto, Ontario is locally known for white squirrel sightings.


Campus populations

In addition to the various towns that boast of their white squirrel populations, a number of university campuses in North America have white squirrels. The
University of Texas at Austin The University of Texas at Austin (UT Austin, UT, or Texas) is a public research university in Austin, Texas. It was founded in 1883 and is the oldest institution in the University of Texas System. With 40,916 undergraduate students, 11,075 ...
is home to a white squirrel population which has spurred the myth of the albino squirrel as a good luck charm. There are many versions of the tale; one of the more popular versions is if one spots the albino squirrel before an exam, they will ace it. The University of North Texas founded the
Albino Squirrel Preservation Society {{unreferenced, date=August 2011 The Albino Squirrel Preservation Society (ASPS) is an international collegiate organization dedicated to "fostering compassion and goodwill" toward albino squirrels. The ASPS has approximately 700 members in eight ...
in 2001, which has since acquired several "worldwide" chapters. In 2006, the University of North Texas held a student referendum to name their white squirrel as the university's secondary mascot, but the vote was narrowly defeated by the student body.
University of Wisconsin - Eau Claire A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, the ...
has a significant white squirrel population both on the campus and in other areas of the city of Eau Claire.
Michigan Technological University Michigan Technological University (Michigan Tech, MTU, or simply Tech) is a public research university in Houghton, Michigan, founded in 1885 as the Michigan Mining School, the first post-secondary institution in the Upper Peninsula of Michiga ...
in Houghton, Michigan is home to frequently sighted white squirrels that live on and around the campus. A
Facebook Facebook is an online social media and social networking service owned by American company Meta Platforms. Founded in 2004 by Mark Zuckerberg with fellow Harvard College students and roommates Eduardo Saverin, Andrew McCollum, Dust ...
group dedicated to these squirrels, called ''I've Seen the Albino Squirrel of Michigan Tech'', was created for people to post photographs and anecdotes of their encounters with the white squirrels, and includes some stories from Michigan Tech alumni that recall seeing white squirrels in Houghton dating back to the 1930s. In Kentucky, the
University of Louisville The University of Louisville (UofL) is a public research university in Louisville, Kentucky. It is part of the Kentucky state university system. When founded in 1798, it was the first city-owned public university in the United States and one ...
has established its own chapter of the Albino Squirrel Preservation Society, which maintains contact with its members and interested parties through a Facebook group by that name. The university has an open policy to give away a free t-shirt to anyone who takes a photograph of a white squirrel on campus grounds and brings it to the administration offices. Other university campuses that have albino squirrel populations include Oberlin College in Ohio,
Ohio State University The Ohio State University, commonly called Ohio State or OSU, is a public land-grant research university in Columbus, Ohio. A member of the University System of Ohio, it has been ranked by major institutional rankings among the best pub ...
in
Columbus, Ohio Columbus () is the state capital and the most populous city in the U.S. state of Ohio. With a 2020 census population of 905,748, it is the 14th-most populous city in the U.S., the second-most populous city in the Midwest, after Chicago, an ...
, Western Kentucky University in Bowling Green, Kentucky (which has had a population of albino squirrels since the 1960s), and Youngstown State University in
Youngstown, Ohio Youngstown is a city in the U.S. state of Ohio, and the largest city and county seat of Mahoning County. At the 2020 census, Youngstown had a city population of 60,068. It is a principal city of the Youngstown–Warren metropolitan area, whi ...
. Michael Stokes, a biology professor at Western Kentucky University, commented that the probable cause for the abundance of white squirrels on university campuses was because they were originally introduced by someone: "We're not sure how they got here, but I'll tell you how it usually happens...When you see them, especially around a college campus or parks, somebody brought them in because they thought it would be neat to have white squirrels around." Albert Meier, another biology professor at Western Kentucky University, added that: "... white squirrels rarely survive in the wild because they can't easily hide. But on a college campus, they are less likely to be consumed by other animals."


In folklore

A story in which a
Nāga The Nagas (IAST: ''nāga''; Devanāgarī: नाग) are a divine, or semi-divine, race of half-human, half-serpent beings that reside in the netherworld (Patala), and can occasionally take human or part-human form, or are so depicted in art. ...
shapeshifts into a white or albino squirrel, is killed by a hunter, and is magically transformed into meat equal to 8,000 cartloads figures prominently in the
folklore Folklore is shared by a particular group of people; it encompasses the traditions common to that culture, subculture or group. This includes oral traditions such as tales, legends, proverbs and jokes. They include material culture, rangin ...
of rocket festival
tradition A tradition is a belief or behavior (folk custom) passed down within a group or society with symbolic meaning or special significance with origins in the past. A component of cultural expressions and folklore, common examples include holidays or ...
s and the origin of Nong Han Kumphawapi Lake in Northeast Thailand.


Red and gray squirrels in the UK

A decline of the
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbe ...
and the rise of the eastern gray squirrel, an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
from North America, has been widely remarked upon in British popular culture. It is mostly regarded as the invading grays driving out the native red species. Evidence also shows that gray squirrels are vectors of the
squirrel parapoxvirus ''Squirrelpox virus (SQPV)'' is a virus that causes the fatal disease squirrelpox in United Kingdom red squirrels. The virus is often carried by grey squirrels from North America, which rarely die from the disease. Elsewhere in the Red Squirrel' ...
for which no vaccine is currently available, and which is deadly to red squirrels but does not seem to affect the non-native host. Currently, the red squirrel's range has been reduced to the
coniferous forests Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ...
in Scotland, and in England's Formby, the
Lake District The Lake District, also known as the Lakes or Lakeland, is a mountainous region in North West England. A popular holiday destination, it is famous for its lakes, forests, and mountains (or '' fells''), and its associations with William Wordswor ...
, Brownsea Island, and the
Isle of Wight The Isle of Wight ( ) is a county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the largest and second-most populous island of England. Referred to as 'The Island' by residents, the Is ...
. The majority of England's red squirrels are found in the county of
Northumberland Northumberland () is a county in Northern England, one of two counties in England which border with Scotland. Notable landmarks in the county include Alnwick Castle, Bamburgh Castle, Hadrian's Wall and Hexham Abbey. It is bordered by land ...
. Special measures are in place to contain and remove any infiltration of gray squirrels into these areas. Though the population has dramatically decreased, they remain listed on the IUCN Red List as Least Concern. As of 2008, the eastern gray squirrel was regarded as
vermin Vermin ( colloquially varmint(s) or varmit(s)) are pests or nuisance animals that spread diseases or destroy crops or livestock. Since the term is defined in relation to human activities, which species are included vary by region and enterp ...
and it was illegal to release any into the wild; any caught could be releases only if one applied for and was granted a licence to do so. As of 2015, any caught in Scotland had to be humanely killed.


See also

*
American red squirrel The American red squirrel (''Tamiasciurus hudsonicus'') is one of three species of tree squirrels currently classified in the genus '' Tamiasciurus'', known as the pine squirrels (the others are the Douglas squirrel, ''T. douglasii'', and the s ...
* Black squirrel * Eastern gray squirrel * Fox squirrel *
Red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbe ...
*
Squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. ...
* Western gray squirrel


References

;Footnotes ;Sources


Further reading

* *Nowak, Ronald M. 1999. ''Walker's Mammals of the World'', 6th edition. Johns Hopkins University Press, 1936 pp., *Young, Miriam. 1964. ''Miss Suzy'', Purple House Press,


External links

{{commons category, Sciurus
The Squirrel Lovers Club
an international club of squirrel enthusiasts.
Squirrel-Rehab.org
includes advice and links for helping orphaned baby squirrels. Squirrels Rodents by common name af:Boomeekhoring