Thomas R. Holtz Jr.
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Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate
palaeontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
, author, and principal lecturer at the
University of Maryland The University of Maryland, College Park (University of Maryland, UMD, or simply Maryland) is a public land-grant research university in College Park, Maryland. Founded in 1856, UMD is the flagship institution of the University System of M ...
's Department of Geology. He has published extensively on the
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...
,
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
,
ecomorphology Ecomorphology or ecological morphology is the study of the relationship between the ecological role of an individual and its morphological adaptations. The term "morphological" here is in the anatomical context. Both the morphology and ecology exh ...
, and locomotion of terrestrial predators, especially on
tyrannosaurid Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera ...
s and other
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s.Holtz, ''Thomas R. Holtz Jr.''. He wrote the book '' Dinosaurs'' and is the author or co-author of the chapters "Saurischia", "Basal Tetanurae", and "Tyrannosauroidea" in the second edition of ''
The Dinosauria ''The Dinosauria'' is an extensive book on dinosaurs, compiled by David B. Weishampel, Peter Dodson, and Halszka Osmólska. It has been published in 2 editions, with the first edition published in 1990, consisting of material from 23 scientists.B ...
''. He has also been consulted as a scientific advisor for the ''
Walking With Dinosaurs ''Walking with Dinosaurs'' is a 1999 six-part nature documentary television miniseries created by Tim Haines and produced by the BBC Science Unit the Discovery Channel and BBC Worldwide, in association with TV Asahi, ProSieben and France 3. Envi ...
'' BBC series as well as the
Discovery Discovery may refer to: * Discovery (observation), observing or finding something unknown * Discovery (fiction), a character's learning something unknown * Discovery (law), a process in courts of law relating to evidence Discovery, The Discover ...
special '' When Dinosaurs Roamed America'', and has appeared in numerous documentaries focused on prehistoric life, such as '' Jurassic Fight Club'' on
History History (derived ) is the systematic study and the documentation of the human activity. The time period of event before the invention of writing systems is considered prehistory. "History" is an umbrella term comprising past events as well ...
and ''
Monsters Resurrected ''Monsters Resurrected'' is an American documentary television series that premiered on September 13, 2009, on the Discovery Channel. The program reconstructs extinct animals of both Mesozoic and Cenozoic. It is also called ''Mega Beasts''. Fe ...
'', '' Dinosaur Revolution'' and '' Clash of the Dinosaurs'' on Discovery. Holtz is also the director of the Science and Global Change Program within the College Park Scholars living-learning community at the University of Maryland, College Park.


Early life

Holtz was born in California. He has stated that he wanted to be a dinosaur as a child, but upon learning that it is impossible to turn into a dinosaur he shifted his goals to study them. He attended
Johns Hopkins University Johns Hopkins University (Johns Hopkins, Hopkins, or JHU) is a private research university in Baltimore, Maryland. Founded in 1876, Johns Hopkins is the oldest research university in the United States and in the western hemisphere. It consi ...
, where he met his future wife. He then attended Yale, where he meet
John Ostrom John Harold Ostrom (February 18, 1928 – July 16, 2005) was an American paleontologist who revolutionized modern understanding of dinosaurs in the 1960s. As first proposed by Thomas Henry Huxley in the 1860s, Ostrom showed that dinosaurs were ...
who served as his academic advisor.


Theories

Holtz has come up with several new theories and hypotheses about the dinosaurs' classification. For example, he coined the terms Maniraptoriformes and Arctometatarsus. He also proposed two classification systems for theropods. The first is the clade Arctometatarsalia, made up of tyrannosauroids, ornithomimosaurs, and
troodontids Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...
, because all of these
coelurosaurs Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...
had pinched middle
metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the me ...
bones in their feet. In this proposed classification system, the tyrannosauroids were supposedly basal to a clade known as Bullatosauria, which was made up of the
Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...
and the
Ornithomimosauria Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and Nor ...
. These two groups were purported to form a clade, because they both shared a common characteristic; which was a skull capsule. However, later on, both of these classification systems were found to be paraphyletic, or "artificial", clades. For example, troodontids are now known to be deinonychosaurs, or "raptors", closely related to dromaeosaurids and birds. The discovery of basal tyrannosauroids, such as ''
Guanlong ''Guanlong'' (冠龍) is a genus of extinct proceratosaurid tyrannosauroid from the Late Jurassic of China. The taxon was first described in 2006 by Xu Xing ''et al.'', who found it to represent a new taxon related to ''Tyrannosaurus''. The na ...
'', which lacked an arctometatarsus, also helped to disprove this theory. And, eventually, the "skull capsule" in troodontids and ornithomimosaurs was found to be an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
, causing the clade Bullatosauria to be abandoned. Holtz was also a key figure in the discovery that tyrannosauroids were not carnosaurs, as had been previously believed by most palaeontologists, but rather large
coelurosaur Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...
s. One of the first scientists to theorize this, Holtz contributed greatly to the debunking of a monophyletic Carnosauria that contained Megalosauroidea, Allosauroidea and Tyrannosauroidea. However, fellow vertebrate paleontologist Oliver Rauhut is known to recover a monophyletic Carnosauria that contains both Megalosauroidea and Allosauroidea in many of his publications, although like most researcher he agrees that Tyrannosauroids are likely Coelurosaurians as based on the conclusions made by Holtz.


Selected publications

* * * * * * *


Footnotes


References

* * {{DEFAULTSORT:Holtz, Thomas R. Jr. Living people American paleontologists University of Maryland, College Park faculty 1965 births