''Tetraclinis'' (also called arar, araar or Sictus tree) is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...

coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ext ...

trees in the cypress family

Cupressaceae Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioeci ...

, containing only one species, ''Tetraclinis articulata'', also known as Thuja articulata,Memidex: sandarac (wood)
Retrieved 2012-05-16 sandarac, sandarac tree or Barbary thuja,

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

to the western

Mediterranean region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and wa ...

. It is native to northwestern Africa in the

Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through ...

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...

Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...

, and

Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...

, with two small outlying populations on

Malta Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...

, and near Cartagena in southeast

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...

. It grows at relatively low altitudes in a hot, dry subtropical

Mediterranean climate A Mediterranean climate (also called a dry summer temperate climate ''Cs'') is a temperate climate sub-type, generally characterized by warm, dry summers and mild, fairly wet winters; these weather conditions are typically experienced in the ...

.Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. Its closest relatives are '' Platycladus,

Microbiota Microbiota are the range of microorganisms that may be commensal, symbiotic, or pathogenic found in and on all multicellular organisms, including plants. Microbiota include bacteria, archaea, protists, fungi, and viruses, and have been found ...

'', and ''

Calocedrus ''Calocedrus'', the incense cedar (alternatively spelled incense-cedar), is a genus of coniferous trees in the cypress family Cupressaceae first described as a genus in 1873. It is native to eastern Asia and western North America. The generic n ...

'', with the closest resemblance to the latter. In older texts, it was sometimes treated in '' Thuja'' or ''

Callitris ''Callitris'' is a genus of coniferous trees in the Cupressaceae (cypress family). There are 16 recognized species in the genus, of which 13 are native to Australia and the other three (''C. neocaledonica, C. sulcata'' and ''C. p ...

'', but it is less closely related to those genera. Alhucemas (11)

It is a small, slow-growing tree, to 6–15 m (rarely 20 m) tall and 0.5 m (rarely 1 m) trunk diameter, often with two or more trunks from the base. The foliage forms in open sprays with scale-like

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

1–8 mm long and 1–1.5 mm broad; the leaves are arranged in opposite decussate pairs, with the successive pairs closely then distantly spaced, so forming apparent whorls of four. The cones are 10–15 mm long, green ripening brown in about 8 months from pollination, and have four thick scales arranged in two opposite pairs. The

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...

s are 5–7 mm long and 2 mm broad, with a 3–4 mm broad papery wing on each side.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins . It is one of only a small number of conifers able to

coppice Coppicing is a traditional method of woodland management which exploits the capacity of many species of trees to put out new shoots from their stump or roots if cut down. In a coppiced wood, which is called a copse, young tree stems are repeate ...

(regrow by sprouting from stumps), an adaptation to survive

wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identi ...

and moderate levels of browsing by animals. Old trees that have sprouted repeatedly over a long period form large burls at the base, known as ''lupias''.

Uses and symbolism

It is the national tree of

, where it is known as ''għargħar'' (derived from the Arabic ). It is now being used locally in afforestation projects. The

resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...

, known as sandarac, is used to make

varnish Varnish is a clear transparent hard protective coating or film. It is not a stain. It usually has a yellowish shade from the manufacturing process and materials used, but it may also be pigmented as desired, and is sold commercially in variou ...

and

lacquer Lacquer is a type of hard and usually shiny coating or finish applied to materials such as wood or metal. It is most often made from resin extracted from trees and waxes and has been in use since antiquity. Asian lacquerware, which may be c ...

; it is particularly valued for preserving

painting Painting is the practice of applying paint, pigment, color or other medium to a solid surface (called the "matrix" or "support"). The medium is commonly applied to the base with a brush, but other implements, such as knives, sponges, and a ...

s. The

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...

, known as thuya wood, citron wood, and alerce, and historically also known as

thyine wood classic drawing of the tree Thyine wood is a 15th-century English name for a wood from the tree known botanically as ''Tetraclinis articulata'' (syn. ''Callitris quadrivalvis'', ''Thuja articulata''). The name is derived from the Greek word ''thuo ...

, is used for decorative woodwork, particularly wood from burls at the base of the trunk. The market in Morocco is unsustainable, focusing as it does on the burl, and has resulted in mass deforestation of the species. The species is also threatened by

overgrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature res ...

, which can kill the coppice regrowth before it gets tall enough to be out of the reach of livestock.

Cultivation

The species is cultivated to be grown as an

ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...

, valued in hot, dry climates. It is also pruned in a

hedge A hedge or hedgerow is a line of closely spaced shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate a road from adjoi ...

form, for privacy and security. The plant can be trained for use as

bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of '' penjing''. Unlike ''penjing'', which utilizes traditional techniques to produc ...

specimens.

Fossil record

†''Tetraclinis salicornioides'' leaf and cone

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of

Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fi ...

age (ca. 5.7 Ma) have been uncovered in Monte Tondo and Borgo Tossignano, northern

Apennines The Apennines or Apennine Mountains (; grc-gre, links=no, Ἀπέννινα ὄρη or Ἀπέννινον ὄρος; la, Appenninus or – a singular with plural meaning;''Apenninus'' (Greek or ) has the form of an adjective, which wou ...

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...

.Palaeoenvironmental analysis of the Messinian macrofossil floras of Tossignano and Monte Tondo (Vena del Gesso Basin, Romagna Apennines, northern Italy) - Vasilis Teodoridis, Zlatko Kvacek, Marco Sami and Edoardo Martinetto - December 2015 DOI: 10.14446/AMNP.2015.249. Immature cones

Gallery

Image:Tetraclinis_articulata8.jpg, ''Tetraclinis articulata'' in the mountains of Cartagena, Spain Image:Araar-root-burl-cup.jpg, A cup made of root burl wood from the Essaouira area of Morocco Image:Tetraclinis_articulata_-_Köhler–s_Medizinal-Pflanzen-270.jpg, Illustration from Koehler's ''Medicinal-Plants'' (1887) File:Tetraclinis articulata MHNT.BOT.2007.52.39.jpg, ''Tetraclinis articulata''

References

{{Taxonbar, from1=Q13091238, from2=Q918555 Cupressaceae Monotypic conifer genera Flora of Tunisia Flora of Algeria Flora of Morocco Flora of Spain Near threatened plants Drought-tolerant plants National symbols of Malta Plants used in bonsai Garden plants of Africa Garden plants of Europe Ornamental trees Drought-tolerant trees