Tazoudasaurus
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''Tazoudasaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of vulcanodontid
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', ' lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s hailing from the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
, located in the
High Atlas High Atlas, also called the Grand Atlas ( ar, الأطلس الكبير, Al-Aṭlas al-Kabīr; french: Haut Atlas; shi, ⴰⴷⵔⴰⵔ ⵏ ⴷⵔⵏ ''Adrar n Dern''), is a mountain range in central Morocco, North Africa, the highest part of t ...
Mountains of
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
in
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
. Along with ''
Ohmdenosaurus ''Ohmdenosaurus'' ("Ohmden lizard") is a genus of sauropod dinosaur that lived during the Early Jurassic epoch in what is now Germany. The only specimen – a tibia (shinbone) and ankle – was discovered in rocks of the Posidonia Shale near H ...
'' is one of the two formally described sauropods from the Toarcian of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
.


Discovery

The remains, consisting of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, a partial adult skeleton (specimen To 2000–1) and associated partial juvenile skeleton (specimen To 2000-2) found in continental detrital sediments of the
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toar ...
aged Azilal Formation, were discovered in the early 2000s and were described by Ronan Allain ''et al.'' in early 2004. The generic name derives from one of the localities, Tazouda, while the specific descriptor is a latinization of the
Berber Berber or Berbers may refer to: Ethnic group * Berbers, an ethnic group native to Northern Africa * Berber languages, a family of Afro-Asiatic languages Places * Berber, Sudan, a town on the Nile People with the surname * Ady Berber (1913–19 ...
term for "slender" due to the
animal Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage ...
's small size for a sauropod. Its fossil was found alongside that of ''
Berberosaurus ''Berberosaurus'' (meaning "Berber lizard", in reference to the Berbers of Morocco) is a genus of neotheropod dinosaur, possibly a ceratosaur, from the Early Toarcian-age (Lower Jurassic) Azilal Formation found in the High Atlas of Toundoute, O ...
'' and an indeterminate large theropod.


Description

''Tazoudasaurus'', a small sauropod at long, is characterized by rather primitive features such as the prosauropod-like mandible with spatulate and denticle-bearing teeth, lack of a U-shaped mandibular
symphysis A symphysis (, pl. symphyses) is a fibrocartilaginous fusion between two bones. It is a type of cartilaginous joint, specifically a secondary cartilaginous joint. # A symphysis is an amphiarthrosis, a slightly movable joint. # A growing togethe ...
as other more derived sauropods. Teeth wear in V-shaped marks indicates tooth occlusion, suggesting that vulcanodontids processed food orally when feeding. The frontal and the parietal are incomplete, the former being broken anteriorly and the latter posteriorly. The neck is flexible with elongate
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e that lack true
pleurocoels Skeletal pneumaticity is the presence of air spaces within bones. It is generally produced during development by excavation of bone by pneumatic diverticula (air sacs) from an air-filled space, such as the lungs or nasal cavity. Pneumatization is h ...
while dorsal and caudal vertebrae series tend to be more rigid. ''T. naimi'' bears the most complete fossil skeleton for Early Jurassic sauropod remains found to date due to the scarcity of exposed strata of that age.


Classification

This sauropod is most closely related to '' Vulcanodon'', differing only in caudal vertebrae features while it also possesses characters that place it outside
Eusauropoda Eusauropoda (meaning "true sauropods") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of ''Shunosaurus'', and possibly ''Barapasau ...
. A cladogram after Pol and colleagues, 2021:


See also

* Atlasaurus *
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through ...


References

{{Taxonbar, from=Q139286 Vulcanodontidae Early Jurassic dinosaurs of Africa Toarcian life Jurassic Morocco Fossils of Morocco Fossil taxa described in 2004 Taxa named by Dale Russell Taxa named by Philippe Taquet