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Armen Leonovich Takhtajan or Takhtajian ( hy, Արմեն Լևոնի Թախտաջյան; russian: Армен Леонович Тахтаджян; surname also transliterated Takhtadjan, Takhtadzhi︠a︡n or Takhtadzhian, pronounced takh-tuh-JAHN; 10 June 1910 – 13 November 2009), was a
Soviet The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, ...
- Armenian
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
, one of the most important figures in 20th century
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae excl ...
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
and
systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic t ...
and
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
. His other interests included morphology of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s,
paleobotany Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeog ...
, and the flora of the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
. He was one of the most influential
taxonomists In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
of the latter twentieth century.


Life


Family

Takhtajan was born in
Shushi / hy, Շուշի , settlement_type = City , image_skyline = ShushaCollection2021.jpg , image_caption = Landmarks of Shusha, from top left:Ghazanchetsots Cathedral • Yukhari Govha ...
, Russian Empire, present-day disputed Nagorno-Karabakh, on 10 June 1910, to a family of Armenian intellectuals. His grandfather Meliksan Takhtadzhyan Petrovich had been born in
Trabzon Trabzon (; Ancient Greek: Tραπεζοῦς (''Trapezous''), Ophitic Pontic Greek: Τραπεζούντα (''Trapezounta''); Georgian: ტრაპიზონი (''Trapizoni'')), historically known as Trebizond in English, is a city on the B ...
, Ottoman Empire and was educated in Italy, on the island of
San Lazzaro degli Armeni San Lazzaro degli Armeni (, "Saint Lazarus of the Armenians"; called Saint Lazarus Island in English sources; hy, Սուրբ Ղազար, Surb Ghazar) is a small island in the Venetian Lagoon which has been home to the monastery of the Mekhita ...
, an Armenian
enclave An enclave is a territory (or a small territory apart of a larger one) that is entirely surrounded by the territory of one other state or entity. Enclaves may also exist within territorial waters. ''Enclave'' is sometimes used improperly to deno ...
, spoke many languages and worked as a journalist. He died in Paris in 1930. His father, Leon Meliksanovich Takhtadzhyan (1884–1950), was born in
Batumi Batumi (; ka, ბათუმი ) is the List of cities and towns in Georgia (country), second largest city of Georgia (country), Georgia and the capital of the Autonomous Republic of Adjara, located on the coast of the Black Sea in Georgia's ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
and was educated as an
agronomist An agriculturist, agriculturalist, agrologist, or agronomist (abbreviated as agr.), is a professional in the science, practice, and management of agriculture and agribusiness. It is a regulated profession in Canada, India, the Philippines, the ...
at
Leipzig University Leipzig University (german: Universität Leipzig), in Leipzig in Saxony, Germany, is one of the world's oldest universities and the second-oldest university (by consecutive years of existence) in Germany. The university was founded on 2 December ...
. Graduating in 1906, he worked on farms in France, Switzerland and the United Kingdom, and made a special study of sheep farming. He became proficient in
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
,
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
,
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ...
, Russian, Georgian and
Azerbaijani Azerbaijani may refer to: * Something of, or related to Azerbaijan * Azerbaijanis * Azerbaijani language See also * Azerbaijan (disambiguation) * Azeri (disambiguation) * Azerbaijani cuisine * Culture of Azerbaijan The culture of Azerbaijan ...
. Arriving in Shushi in 1908, then a centre of sheep farming in the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
, looking for work, Leon was forced to teach German at the local ''
Realschule ''Realschule'' () is a type of secondary school in Germany, Switzerland and Liechtenstein. It has also existed in Croatia (''realna gimnazija''), the Austrian Empire, the German Empire, Denmark and Norway (''realskole''), Sweden (''realskola''), ...
'' attached to the Armenian
seminary A seminary, school of theology, theological seminary, or divinity school is an educational institution for educating students (sometimes called ''seminarians'') in scripture, theology, generally to prepare them for ordination to serve as clergy ...
, due to lack of opportunities in his chosen field. There he met and married Gerseliya Sergeevna Gazarbekyan (1887–1974), Armen Takhtajan's mother, a native of Susha, in 1909.


Early life and education

The Takhtajans had three children, Armen (1910–2009), Nellie (1914–1994) and Nora (1918–1965). In 1918 the family were forced to flee to northern Armenia because of the
pogroms A pogrom () is a violent riot incited with the aim of massacring or expelling an ethnic or religious group, particularly Jews. The term entered the English language from Russian to describe 19th- and 20th-century attacks on Jews in the Russian ...
. Throughout his childhood, Armen showed a keen interest in natural history, travelling with his father. Armen attended school in
Tbilisi Tbilisi ( ; ka, თბილისი ), in some languages still known by its pre-1936 name Tiflis ( ), is the capital and the largest city of Georgia, lying on the banks of the Kura River with a population of approximately 1.5 million pe ...
in nearby Georgia, initially at Unified Labor School number 42 (former Mantashevskom commercial school). There he came under the influence of one of his teachers, Alexander Konstantinovich Makaev (Makashvili) (1896–1962), who had previously taught agriculture at Tbilisi State University, and had produced a dictionary of botanical names in Georgian, Russian and Latin. Makaev would take Armen on botanical excursions, teaching him to identify plants from Sosnowski and Grossheim's "Determinants of plant life in the vicinity of Tbilisi" (1920). In 1928 he completed secondary school and travelled to
Leningrad Saint Petersburg ( rus, links=no, Санкт-Петербург, a=Ru-Sankt Peterburg Leningrad Petrograd Piter.ogg, r=Sankt-Peterburg, p=ˈsankt pʲɪtʲɪrˈburk), formerly known as Petrograd (1914–1924) and later Leningrad (1924–1991), i ...
. There he volunteered at the biology school at
Leningrad University Saint Petersburg State University (SPBU; russian: Санкт-Петербургский государственный университет) is a public research university in Saint Petersburg, Russia. Founded in 1724 by a decree of Peter the G ...
and attended lectures by
Vladimir Leontyevich Komarov Vladimir Leontyevich Komarov (russian: Влади́мир Лео́нтьевич Комаро́в; – 5 December 1945) was a Russian and Soviet botanist. Biography Komarov was born in 1869. He was a graduate of St. Petersbur ...
(1869–1945) on
plant morphology Phytomorphology is the study of the physical form and external structure of plants.Raven, P. H., R. F. Evert, & S. E. Eichhorn. ''Biology of Plants'', 7th ed., page 9. (New York: W. H. Freeman, 2005). . This is usually considered distinct from ...
. In 1929 he began his studies in biology at
Yerevan State University Yerevan State University (YSU; hy, Երևանի Պետական Համալսարան, ԵՊՀ, ''Yerevani Petakan Hamalsaran''), also simply University of Yerevan, is the oldest continuously operating public university in Armenia. Founded in 1919 ...
in
Yerevan Yerevan ( , , hy, Երևան , sometimes spelled Erevan) is the capital and largest city of Armenia and one of the world's oldest continuously inhabited cities. Situated along the Hrazdan River, Yerevan is the administrative, cultural, and i ...
, Armenia, which he completed in 1931. He then returned to Tbilisi, enrolling in the All-Union Institute of Subtropical Crops. In 1932 after completing his course at Tbilisi he worked for a while as a laboratory assistant at
Sukhumi Sukhumi (russian: Суху́м(и), ) or Sokhumi ( ka, სოხუმი, ), also known by its Abkhaz name Aqwa ( ab, Аҟәа, ''Aqwa''), is a city in a wide bay on the Black Sea's eastern coast. It is both the capital and largest city of ...
, Georgia, at the subtropical branch of the All-Union Institute of Applied Botany and New Crops (now the
Institute of Plant Industry The Institute of Plant Industry, Vavilov Institute of Plant Industry or All-Russian Research Institute of Plant Industry (in russian: Всероссийский институт растениеводства им. Н. И. Вавилова), as ...
), before returning to Yerevan. In Yerevan he took a position as researcher at the Natural History Museum of Armenia, and then at the Herbarium of the Armenian branch of the Institute of Biology, Soviet Academy of Sciences, and began teaching at Yerevan University in 1936, while completing his Master's thesis. He died in Saint Petersburg on 13 November 2009, at the age of 99, in 2009, having just completed his most important work, ''Flowering Plants''.


Work

From 1938 to 1948 he headed a Department at the
Yerevan State University Yerevan State University (YSU; hy, Երևանի Պետական Համալսարան, ԵՊՀ, ''Yerevani Petakan Hamalsaran''), also simply University of Yerevan, is the oldest continuously operating public university in Armenia. Founded in 1919 ...
, and in 1944–1948 he was director of the Botanical Institute of the Academy of Sciences of the Armenian SSR, and Professor of the
Leningrad State University Saint Petersburg State University (SPBU; russian: Санкт-Петербургский государственный университет) is a public research university in Saint Petersburg, Russia. Founded in 1724 by a decree of Peter the G ...
. Takhtajan was a member of the
Russian Academy of Sciences The Russian Academy of Sciences (RAS; russian: Росси́йская акаде́мия нау́к (РАН) ''Rossíyskaya akadémiya naúk'') consists of the national academy of Russia; a network of scientific research institutes from across t ...
, as well as a foreign associate of the U.S.
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nat ...
since 1971. He was also the academician of the Academy of Sciences of the Armenian SSR, the president of the Soviet All-Union Botanical Society (1973) and the International Association for Plant Taxonomy (1975), member of the Finnish Academy of Science and Literature (1971), the German Academy of Naturalists "Leopoldina" (1972) and other scientific societies. He was a member of the
Norwegian Academy of Science and Letters The Norwegian Academy of Science and Letters ( no, Det Norske Videnskaps-Akademi, DNVA) is a learned society based in Oslo, Norway. Its purpose is to support the advancement of science and scholarship in Norway. History The Royal Frederick Unive ...
from 1980. While at the
Komarov Botanical Institute The Komarov Botanical Institute of the Russian Academy of Sciences (russian: Ботанический институт им. В.Л.Комарова РАН) is a leading botanical institution in Russia, It is located on Aptekarsky Island in St. Pete ...
in
Leningrad Saint Petersburg ( rus, links=no, Санкт-Петербург, a=Ru-Sankt Peterburg Leningrad Petrograd Piter.ogg, r=Sankt-Peterburg, p=ˈsankt pʲɪtʲɪrˈburk), formerly known as Petrograd (1914–1924) and later Leningrad (1924–1991), i ...
in 1940, Takhtajan developed his classification scheme for
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s, which emphasized
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
relationships between plants. His system did not become known to botanists in the
West West or Occident is one of the four cardinal directions or points of the compass. It is the opposite direction from east and is the direction in which the Sun sets on the Earth. Etymology The word "west" is a Germanic word passed into some ...
until after 1950, and in the late 1950s he began a correspondence and collaboration with the prominent American botanist
Arthur Cronquist Arthur John Cronquist (March 19, 1919 – March 22, 1992) was an American biologist, botanist and a specialist on Compositae. He is considered one of the most influential botanists of the 20th century, largely due to his formulation of the Cr ...
, whose plant classification scheme was heavily influenced by his collaboration with Takhtajan and other botanists at Komarov. He is chiefly famous as the author of works on the origins of flowering plants and paleobotany, developing a new classification system of higher plants. He worked on the "Flora of Armenia" (vol. 1–6, 1954–73) and "Fossil flowering plants of the USSR "(v. 1, 1974) books. Takhtajan also developed a system of
floristic region A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
s.(Takhtajan, Crovello and Cronquist, 1986) For many years restrictions were placed on his work because of his opposition to the official line on genetics promoted by Lysenko. In 1993 he worked for a while at the
New York Botanical Garden The New York Botanical Garden (NYBG) is a botanical garden at Bronx Park in the Bronx, New York City. Established in 1891, it is located on a site that contains a landscape with over one million living plants; the Enid A. Haupt Conservatory, ...
.


Takhtajan system

The "
Takhtajan system A system of plant taxonomy, the Takhtajan system of plant classification was published by Armen Takhtajan, in several versions from the 1950s onwards. It is usually compared to the Cronquist system. It admits paraphyletic groups. Systems The f ...
" of flowering plant classification treats flowering plants as a
division Division or divider may refer to: Mathematics *Division (mathematics), the inverse of multiplication *Division algorithm, a method for computing the result of mathematical division Military *Division (military), a formation typically consisting ...
(phylum), ''
Magnoliophyta Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
'', with two classes, ''
Magnoliopsida Magnoliopsida is a valid botanical name for a class of flowering plants. By definition the class will include the family Magnoliaceae, but its circumscription can otherwise vary, being more inclusive or less inclusive depending upon the classif ...
'' (dicots) and ''
Liliopsida Liliopsida Batsch (synonym: Liliatae) is a botanical name for the class containing the family Liliaceae (or Lily Family). It is considered synonymous (or nearly synonymous) with the name monocotyledon. Publication of the name is credited to Scopol ...
'' (monocots). These two classes are subdivided into subclasses, and then superorders, orders, and families. The Takhtajan system is similar to the
Cronquist system The Cronquist system is a taxonomic classification system of flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of Flowering Plants'' (1968; 2nd edition, 1988) ...
, but with somewhat greater complexity at the higher levels. He favors smaller orders and families, to allow character and evolutionary relationships to be more easily grasped. The Takhtajan classification system remains influential; it is used, for example, by the Montréal Botanical Garden.


Selected publications

* 1948. Морфологическая эволюция покрытосеменных (Morphological evolution of angiosperms). ranslated from Russian to German by Werner Höppner, as ''Die Evolution der Angiospermae'' 1959, Fischer, Jena * * 1969. Flowering plants: origin and dispersal. Oliver and Boyd, Edinburgh ranslated from Russian by C Jeffreyref name=T1966/> * 2009. Flowering Plants. Springer, New York


Legacy

Takhtajan has been considered one of the leading botanists of his time. He has been honoured in the naming of several plant genera. In 1980, botanist Vandika Ervandovna Avetisyan published '' Takhtajaniella'', which is a genus of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
from Transcausica, belonging to the family
Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some are shrubs. The leav ...
and it was named in his honour. Then in 1990, Nazarova published '' Takhtajaniantha'' (from the dandelion tribe within the
daisy family The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
,Nazarova, Estella A. 1990. Biologicheskii Zhurnal Armenii 43: 179–183) and lastly in 1997, ''
Takhtajania ''Takhtajania'' is a genus of flowering plants of the family Winteraceae, which contains a single species, ''Takhtajania perrieri''. It is endemic to Madagascar. ''Takhtajania'' is found in a small area of the Madagascar subhumid forests. It is ...
'' (from the family
Winteraceae Winteraceae is a primitive family of tropical trees and shrubs including 93 species in five genera. It is of particular interest because it is such a primitive angiosperm family, distantly related to Magnoliaceae, though it has a much more s ...
, which was found in
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
) was published.World Conservation Monitoring Centre. 1998
''Takhtajania perrieri''.
The IUCN Red List of Threatened Species 1998. Downloaded on 10 October 2015.


See also

*
Takhtajan system A system of plant taxonomy, the Takhtajan system of plant classification was published by Armen Takhtajan, in several versions from the 1950s onwards. It is usually compared to the Cronquist system. It admits paraphyletic groups. Systems The f ...


References


Works by Takhtajan

* * *Takhtajan, Armen. 1973. Chetyre tsarstva organicheskogo mira (Four Kingdoms of the Organic World). Priroda (Nature), Akad. Nauk 2: 22–32. * * * * * *


External links

*
The parting with Armen Takhtajan
Photo-report on the site of the Komarov Botanical Institute.

{{DEFAULTSORT:Takhtajan, Armen 1910 births 2009 deaths Russian people of Armenian descent Armenian botanists Full Members of the USSR Academy of Sciences Full Members of the Russian Academy of Sciences Phytogeographers Pteridologists Soviet botanists Scientists from Shusha Symbiogenesis researchers Foreign associates of the National Academy of Sciences Members of the German Academy of Sciences Leopoldina Members of the Norwegian Academy of Science and Letters Yerevan State University alumni Academics of Yerevan State University