The littoral zone or nearshore is the part of a

sea The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...

lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...

, or

river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of ...

that is close to the

shore A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past a ...

. In coastal ecology, the littoral zone includes the

intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...

extending from the

high water mark A high water mark is a point that represents the maximum rise of a body of water over land. Such a mark is often the result of a flood, but high water marks may reflect an all-time high, an annual high (highest level to which water rose tha ...

(which is rarely inundated), to coastal areas that are permanently submerged — known as the ''foreshore'' — and the terms are often used interchangeably. However, the geographical meaning of ''littoral zone'' extends well beyond the intertidal zone to include all neritic waters within the bounds of

continental shelves A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

Etymology

The word ''littoral'' may be used both as a

noun A noun () is a word that generally functions as the name of a specific object or set of objects, such as living creatures, places, actions, qualities, states of existence, or ideas.Example nouns for: * Living creatures (including people, alive, ...

and as an

adjective In linguistics, an adjective ( abbreviated ) is a word that generally modifies a noun or noun phrase or describes its referent. Its semantic role is to change information given by the noun. Traditionally, adjectives were considered one of the ...

. It derives from the

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

noun ''litus, litoris'', meaning "shore". (The doubled ''t'' is a late-medieval innovation, and the word is sometimes seen in the more classical-looking spelling ''litoral''.)

Description

The term has no single definition. What is regarded as the full extent of the littoral zone, and the way the littoral zone is divided into subregions, varies in different contexts. For lakes, the littoral zone is the nearshore habitat where

photosynthetically active radiation Photosynthetically active radiation (PAR) designates the spectral range (wave band) of solar radiation from 400 to 700 nanometers that photosynthetic organisms are able to use in the process of photosynthesis. This spectral region corresponds more o ...

penetrates to the lake bottom in sufficient quantities to support photosynthesis. The use of the term also varies from one part of the world to another, and between different disciplines. For example, military commanders speak of the littoral in ways that are quite different from the definition used by marine biologists. The adjacency of water gives a number of distinctive characteristics to littoral regions. The erosive power of water results in particular types of

landform A landform is a natural or anthropogenic land feature on the solid surface of the Earth or other planetary body. Landforms together make up a given terrain, and their arrangement in the landscape is known as topography. Landforms include hills, ...

s, such as

sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...

s, and

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...

. The natural movement of the littoral along the coast is called the littoral drift. Biologically, the ready availability of water enables a greater variety of plant and animal life, and particularly the formation of extensive

wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The p ...

. In addition, the additional local

humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...

due to evaporation usually creates a

microclimate A microclimate (or micro-climate) is a local set of atmospheric conditions that differ from those in the surrounding areas, often with a slight difference but sometimes with a substantial one. The term may refer to areas as small as a few squ ...

supporting unique types of organisms.

In oceanography and marine biology

oceanography Oceanography (), also known as oceanology and ocean science, is the scientific study of the oceans. It is an Earth science, which covers a wide range of topics, including ecosystem dynamics; ocean currents, waves, and geophysical fluid dynami ...

and

marine biology Marine biology is the scientific study of the biology of marine life, organisms in the sea. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology classifi ...

, the idea of the littoral zone is extended roughly to the edge of the

continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

. Starting from the shoreline, the littoral zone begins at the spray region just above the high tide mark. From here, it moves to the intertidal region between the high and low water marks, and then out as far as the edge of the

. These three subregions are called, in order, the supralittoral zone, the eulittoral zone, and the sublittoral zone.

Supralittoral zone

The ''supralittoral zone'' (also called the ''splash'', ''spray'' or ''supratidal zone'') is the area above the spring high tide line that is regularly splashed, but not submerged by ocean water. Seawater penetrates these elevated areas only during storms with high tides. Organisms that live here must cope with exposure to fresh water from rain, cold, heat, dryness and

predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

by land animals and seabirds. At the top of this area, patches of dark

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.periwinkles, Neritidae and detritus feeding

Isopod Isopoda is an order of crustaceans that includes woodlice and their relatives. Isopods live in the sea, in fresh water, or on land. All have rigid, segmented exoskeletons, two pairs of antennae, seven pairs of jointed limbs on the thorax, and ...

a commonly inhabit the lower supralittoral.Yip and Madl

Eulittoral zone

The ''eulittoral zone'' (also called the ''midlittoral'' or ''mediolittoral zone'') is the ''intertidal zone'', known also as the ''foreshore''. It extends from the spring high tide line, which is rarely inundated, to the spring low tide line, which is rarely not inundated. It is alternately exposed and submerged once or twice daily. Organisms living here must be able to withstand the varying conditions of temperature, light, and salinity. Despite this,

productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...

is high in this zone. The wave action and turbulence of recurring tides shape and reform cliffs, gaps and caves, offering a huge range of habitats for sedentary organisms. Protected rocky shorelines usually show a narrow, almost homogenous, eulittoral strip, often marked by the presence of

barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosiv ...

s. Exposed sites show a wider extension and are often divided into further zones. For more on this, see

intertidal ecology Intertidal ecology is the study of intertidal ecosystems, where organisms live between the low and high tide lines. At low tide, the intertidal is exposed whereas at high tide, the intertidal is underwater. Intertidal ecologists therefore study th ...

Sublittoral zone

The ''sublittoral zone'' starts immediately below the eulittoral zone. This zone is permanently covered with seawater and is approximately equivalent to the

neritic zone The neritic zone (or sublittoral zone) is the relatively shallow part of the ocean above the drop-off of the continental shelf, approximately in depth. From the point of view of marine biology it forms a relatively stable and well-illuminated ...

. In

physical oceanography Physical oceanography is the study of physical conditions and physical processes within the ocean, especially the motions and physical properties of ocean waters. Physical oceanography is one of several sub-domains into which oceanography is div ...

, the sublittoral zone refers to coastal regions with significant tidal flows and energy dissipation, including non-linear flows, internal waves, river outflows and oceanic fronts. In practice, this typically extends to the edge of the

, with depths around 200 meters. In marine biology, the sublittoral zone refers to the areas where sunlight reaches the ocean floor, that is, where the water is never so deep as to take it out of the

photic zone The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...

. This results in high

primary production In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...

and makes the sublittoral zone the location of the majority of sea life. As in physical oceanography, this zone typically extends to the edge of the

. The benthic zone in the sublittoral is much more stable than in the intertidal zone; temperature, water pressure, and the amount of sunlight remain fairly constant. Sublittoral corals do not have to deal with as much change as intertidal corals.

Coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...

s can live in both zones, but they are more common in the sublittoral zone. Within the sublittoral, marine biologists also identify the following: *The infralittoral zone is the algal dominated zone, which may extend to five metres below the low water mark. *The circalittoral zone is the region beyond the infralittoral, that is, below the algal zone and dominated by sessile animals such as

mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which ...

s and

oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not ...

s. Shallower regions of the sublittoral zone, extending not far from the shore, are sometimes referred to as the subtidal zone.

Habitats in littoral zones

Many vertebrates (e.g., mammals, waterfowl, reptiles) and invertebrates (insects, etc.) use both the littoral zone as well as the terrestrial ecosystem for food and habitat. Biota that are commonly assumed to reside in the

pelagic zone The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...

often rely heavily on resources from the littoral zone. Littoral areas of ponds and lakes are typically better oxygenated, structurally more complex, and afford more abundant and diverse food resources than do profundal sediments. All these factors lead to a high diversity of insects and very complex trophic interactions. The great lakes of the world represent a global heritage of surface freshwater and aquatic biodiversity. Species lists for 14 of the world’s largest lakes reveal that 15% of the global diversity (the total number of species) of freshwater fishes, 9% of non-insect freshwater invertebrate diversity, and 2% of aquatic insect diversity live in this handful of lakes. The vast majority (more than 93%) of species inhabit the shallow, nearshore littoral zone, and 72% are completely restricted to the littoral zone, even though littoral habitats are a small fraction of total lake areas. Because the littoral zone is important for many recreational and industrial purposes, it is often severely affected by many human activities that increase nutrient loading, spread invasive species, cause acidification and climate change, and produce increased fluctuations in water level. Littoral zones are both more negatively affected by human activity and less intensively studied than offshore waters. Conservation of the remarkable biodiversity and biotic integrity of large lakes will require better integration of littoral zones into our understanding of lake ecosystem functioning and focused efforts to alleviate human impacts along the shoreline.

In freshwater ecosystems

In freshwater situations, the littoral zone is the nearshore habitat where

penetrates to the lake bottom in sufficient quantities to support photosynthesis. Sometimes other definitions are used. For example, the

Minnesota Department of Natural Resources The Minnesota Department of Natural Resources, or Minnesota DNR, is the agency of the U.S. state of Minnesota charged with conserving and managing the state's natural resources. The agency maintains areas such as state parks, state forests, rec ...

defines littoral as that portion of the lake that is less than 15 feet in depth. Such fixed-depth definitions often do not accurately represent the true ecological zonation, but are sometimes used because they are simple measurements to make bathymetric maps or when there are no measurements of light penetration. The littoral zone comprises an estimated 78% of Earth's total lake area. The littoral zone may form a narrow or broad fringing wetland, with extensive areas of aquatic plants sorted by their tolerance to different water depths. Typically, four zones are recognized, from higher to lower on the shore: wooded wetland,

wet meadow A wet meadow is a type of wetland with soils that are Solubility, saturated for part or all of the growing season. Debate exists whether a wet meadow is a type of marsh or a completely separate type of wetland. Wet prairies and wet savannas are hyd ...

marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found ...

and aquatic vegetation.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. Chapter 2. The relative areas of these four types depends not only on the profile of the shoreline, but upon past water levels. The area of wet meadow is particularly dependent upon past water levels;Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. in general, the area of wet meadows along lakes and rivers increases with natural water level fluctuations. Many of the animals in lakes and rivers are dependent upon the wetlands of littoral zones, since the rooted plants provide habitat and food. Hence, a large and productive littoral zone is considered an important characteristic of a healthy lake or river. Littoral zones are at particular risk for two reasons. First, human settlement is often attracted to shorelines, and settlement often disrupts breeding habitats for littoral zone species. For example, many turtles are killed on roads when they leave the water to lay their eggs in upland sites. Fish can be negatively affected by docks and retaining walls which remove breeding habitat in shallow water. Some shoreline communities even deliberately try to remove wetlands since they may interfere with activities like swimming. Overall, the presence of human settlement has a demonstrated negative impact upon adjoining wetlands. An equally serious problem is the tendency to stabilize lake or river levels with dams. Dams removed the spring flood, which carries nutrients into littoral zones and reduces the natural fluctuation of water levels upon which many wetland plants and animals depend. Hence, over time, dams can reduce the area of wetland from a broad littoral zone to a narrow band of vegetation. Marshes and wet meadows are at particular risk.

Other definitions

For the purposes of naval operations, the

US Navy The United States Navy (USN) is the maritime service branch of the United States Armed Forces and one of the eight uniformed services of the United States. It is the largest and most powerful navy in the world, with the estimated tonnage ...

divides the littoral zone in the ways shown on the diagram at the top of this article. The

US Army Corps of Engineers , colors = , anniversaries = 16 June (Organization Day) , battles = , battles_label = Wars , website = , commander1 = ...

and the US Environmental Protection Agency have their own definitions, which have legal implications. The UK

Ministry of Defence {{unsourced, date=February 2021 A ministry of defence or defense (see spelling differences), also known as a department of defence or defense, is an often-used name for the part of a government responsible for matters of defence, found in state ...

defines the littoral as ''those land areas (and their adjacent areas and associated air space) that are susceptible to engagement and influence from the sea'',.

References

Sources

* Haslett, Simon K (2001) ''Coastal Systems.'' Routledge. * Mann, Kenneth Henry (2000
''Ecology of Coastal Waters''
Blackwell Publishing. * Yip, Maricela and Madl, Pierre (1999

University of Salzburg The University of Salzburg (german: Universität Salzburg), also known as the Paris Lodron University of Salzburg (''Paris-Lodron-Universität Salzburg'', PLUS), is an Austrian public university in Salzburg, Salzburg municipality, Salzburg (state ...

. {{DEFAULTSORT:Littoral Zone Aquatic biomes Marine biology Aquatic ecology Habitats Coasts Fisheries science Coastal geography Oceanographical terminology Limnology Oceanography