Spirostachys africana
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''Spirostachys africana'' is a medium-sized (about tall) deciduous
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
with a straight, clear trunk, occurring in the warmer parts of
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
. Its wood is known as tamboti, tambotie, tambootie or tambuti. It prefers growing in single-species
copse Coppicing is a traditional method of woodland management which exploits the capacity of many species of trees to put out new shoots from their stump or roots if cut down. In a coppiced wood, which is called a copse, young tree stems are repeate ...
s in deciduous woodland, often along watercourses or on brackish flats and sandy soils.


Description

The leaves are small, elliptic with crenate margins, and turn bright red in winter before dropping. The petiole has 2 small glands at the distal end. The grey-black rough
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
is distinctively split into neat rectangles. The
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
-like
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s appear in early spring before the leaves. Male and female flowers are borne separately on the same tree (
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is ...
). The small 3-lobed capsules or
schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...
s split into three equal
indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...
segments ( mericarps or
cocci A coccus (plural cocci) is any bacterium or archaeon that has a spherical, ovoid, or generally round shape. Bacteria are categorized based on their shapes into three classes: cocci (spherical-shaped), bacillus (rod-shaped) and spiral ( of whi ...
) when ripe; on a warm day this splitting ( dehiscence) can sound like a distant
fusillade A fusillade is the simultaneous and continuous firing of a group of firearms on command. It stems from the French word ''fusil'', meaning firearm, and ''fusiller'' meaning to shoot. In the context of military tactics, the term is generally used t ...
of shots. The seeds are globose with a chartaceous testa.


Wood and toxicity

Despite it being prone to
heart-rot 300px, The bracket fungus '' Fistulina hepatica'' is one of many that cause heart rot. In trees, heart rot is a fungal disease that causes the decay of wood at the center of the trunk and branches. Fungi enter the tree through wounds in the bark ...
, it is prized in the
furniture Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating ( tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks) ...
industry for its beautiful, dense and durable
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
, which is reddish-brown with darker streaks, a satin-like lustre and extremely fragrant sweet, spicy smell. The underbark exudes a white, poisonous
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...
when freshly cut, and campfires that burn tamboti fuel give off noxious fumes contaminating meat or other food grilled on the open flames or coals. The latex is used as a
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
poison, is applied to arrow-tips and is used as a
purgative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...
by indigenous tribes.


Jumping beans

The fruits while green are frequently parasitised by the small grey
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
'' Emporia melanobasis'' (
Pyralidae The Pyralidae, commonly called pyralid moths, snout moths or grass moths, are a family of Lepidoptera in the ditrysian superfamily Pyraloidea. In many (particularly older) classifications, the grass moths (Crambidae) are included in the Pyralida ...
: Phycitinae). Larvae develop within the growing fruits which show no external damage. When the fruits are mature each splits into 3 cocci. The larvae jack-knife inside the fallen segments, causing them to move about erratically and vigorously, to the surprise of the uninitiated. This has led to the name "jumping bean tree". The
Mexican jumping bean Mexican jumping beans (also known as ' in Spanish) are seed pods that have been inhabited by the larva of a small moth (''Cydia saltitans'') and are native to Mexico. The "bean" is usually tan to brown. They are from the shrub '' Sebastiania pa ...
, ''
Sebastiania ''Sebastiania'' is a genus of flowering plants in the family Euphorbiaceae first described in 1821. It is native to North and South America from Arizona and the West Indies south to Uruguay.Webster, G. L. & M.J. Huft. 1988. Revised synopsis ...
sp.'', also belongs to the family
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as '' Euphorbia paralias'', are herbs, but some, ...
and is parasitised by the moth ''
Cydia saltitans ''Cydia saltitans'' or jumping bean moth is a moth from Mexico that is most widely known as its larva, where it inhabits the carpels of seeds from several related shrubby trees, mainly '' Sebastiania pavoniana'' or '' Sapium biloculare'' (syn. ...
''.


Gallery

Spirostachys africana, lower, Skrikfontein, a.jpg, ''S. africana'' foliage File:Spirostachys africana capsule.JPG, fruit capsule or
schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...
Spiafr01.jpg, fruit and fruit segments or mericarps Spiafr03.jpg, mericarp with ''Emporia melanobasis'' larva Spiafr12a.jpg, Wood Spiafr05a.jpg, Log with heart-rot


See also

*
List of Southern African indigenous trees This is a list of Southern African trees, shrubs, suffrutices, geoxyles and lianes, and is intended to cover Angola, Botswana, Lesotho, Malawi, Mozambique, Namibia, South Africa, Swaziland, Zambia and Zimbabwe. The notion of 'indigenous' is of ...


References

*''Medicinal and Poisonous Plants of Southern and Eastern Africa'' - John Mitchell Watt & Maria Gerdina Breyer-Brandwyk (E&S Livingstone 1962) * {{Taxonbar, from=Q7578203 Hippomaneae Trees of South Africa Flora of Mozambique Taxa named by Otto Wilhelm Sonder