''Spinops'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

centrosaurine Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day A ...

ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurass ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest T ...

, southern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...

. It was a medium-sized ceratopsian, reaching in length and in body mass.

Discovery and naming

''Spinops'' is known from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

NHMUK R16307, a partial parietal bone, preserving most of the midline bar. Referred material include NHMUK R16308, a partial parietal bone, partial

dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

and unidentifiable

limb Limb may refer to: Science and technology * Limb (anatomy), an appendage of a human or animal *Limb, a large or main branch of a tree *Limb, in astronomy, the curved edge of the apparent disk of a celestial body, e.g. lunar limb *Limb, in botany, ...

fragments, NHMUK R16306, an incomplete

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...

, preserving only the

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...

portion of the skull, and NHMUK R16309, a partial right squamosal. None of this material was found in articulation, however it was all closely associated in the same bone bed, in the northwestern region ("Steveville Badlands") of the

Dinosaur Provincial Park Dinosaur Provincial Park is a UNESCO World Heritage Site situated a two hour drive east of Calgary, Alberta, Canada; or , about a half-hour drive northeast of Brooks. The park is situated in the Red Deer River valley, which is noted for its stri ...

. Fossils of ''Spinops'' were first found in 1916, and were housed in the

Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history collections that include current and historical records of animals, plants, fungi, ecosystems, geology, paleontology, climatology, and more. ...

in London. The material was not described until 2011, when the new species ''Spinops sternbergorum'' was erected. The material was probably collected from the upper part of the

Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...

or the lower part of the

Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...

, dating to the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...

stage of the

period. Two partials skulls of ''Spinops'' were found in 1916, in a large bone bed near the

Red Deer River The Red Deer River is a river in Alberta and a small portion of Saskatchewan, Canada. It is a major tributary of the South Saskatchewan River and is part of the larger Saskatchewan-Nelson system that empties into Hudson Bay. Red Deer River h ...

of southern Alberta, by American commercial fossil collector

Charles Hazelius Sternberg Charles Hazelius Sternberg (June 15, 1850 – July 20, 1943) was an American fossil collector and paleontologist. He was active in both fields from 1876 to 1928, and collected fossils for Edward Drinker Cope and Othniel C. Marsh, and for the ...

and his son Levi Sternberg. The fossils were sent to the Natural History Museum in London (then called the British Museum (Natural History)), which had financed the expedition. The museum considered the fossils too fragmentary to display, leaving them unprepared in the collections. In a letter to Charles H. Sternberg, English paleontologist

Arthur Smith Woodward Sir Arthur Smith Woodward, FRS (23 May 1864 – 2 September 1944) was an English palaeontologist, known as a world expert in fossil fish. He also described the Piltdown Man fossils, which were later determined to be fraudulent. He is not relate ...

of the British Museum called the ''Spinops'' material "nothing but rubbish". The precise whereabouts of the bonebed that yielded the fossils is unknown due to poor field record keeping, but

Darren Tanke Darren H. Tanke (born 1960) is a Canadian fossil preparation technician of the Dinosaur Research Program at the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta. Born in Calgary, Tanke became interested in natural history at an early ...

of the

Royal Tyrrell Museum The Royal Tyrrell Museum of Palaeontology (RTMP, and often referred to as the Royal Tyrrell Museum) is a palaeontology museum and research facility in Drumheller, Alberta, Canada. The museum was named in honour of Joseph Burr Tyrrell, and is situ ...

is spearheading attempts at its relocation. The fossils were re-examined in 2011 by a team led by Dr Andrew A. Farke; which realized that the fossils represented an entirely new species of dinosaur. ''Spinops'' was first named by Andrew A. Farke, Michael J. Ryan, Paul M. Barrett,

Darren H. Tanke Darren is a masculine given name of uncertain etymological origins. Some theories state that it originated from an Anglicisation of the Irish first name Darragh or Dáire, meaning "Oak Tree". According to other sources, it is thought to come from ...

, Dennis R. Braman, Mark A. Loewen, and Mark R. Graham in

2011 File:2011 Events Collage.png, From top left, clockwise: a protester partaking in Occupy Wall Street heralds the beginning of the Occupy movement; protests against Libyan dictator Muammar Gaddafi, who was killed that October; a young man celebrates ...

; the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

is ''Spinops sternbergorum''. The generic name is derived from

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

''spina'', "spine", and

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''ops'', "face", in reference to the unique ornamentation on the face. The specific name honours Charles H. and Levi Sternberg.

Classification

The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

presented below follows a recent phylogenetic analysis by Chiba ''et al.'' (2017):

References

{{Taxonbar, from=Q764899 Centrosaurines Late Cretaceous dinosaurs of North America Fossil taxa described in 2011 Paleontology in Alberta Campanian genus first appearances Campanian genus extinctions Ornithischian genera

Discovery and naming

Classification

See also

References