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botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, a bract is a modified or specialized
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
, especially one associated with a reproductive structure such as a
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
,
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
axis or
cone scale A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...
. Bracts are usually different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture. Typically, they also look different from the parts of the flower, such as the petals or sepals. A plant having bracts is referred to as bracteate or bracteolate, while one that lacks them is referred to as ebracteate and ebracteolate, without bracts.


Variants

Some bracts are brightly-coloured and serve the function of attracting pollinators, either together with the
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla ( petals) or tepals when ...
or instead of it. Examples of this type of bract include those of ''
Euphorbia pulcherrima The poinsettia ( or ) (''Euphorbia pulcherrima'') is a commercially important flowering plant species of the diverse spurge family Euphorbiaceae. Indigenous to Mexico and Central America, the poinsettia was first described by Europeans in 1834 ...
'' (poinsettia) and ''
Bougainvillea ''Bougainvillea'' ( , ) is a genus of thorny ornamental vines, bushes, and trees belonging to the four o' clock family, Nyctaginaceae. It is native to eastern South America, found from Brazil, west to Peru, and south to southern Argentina. ...
'': both of these have large colourful bracts surrounding much smaller, less colourful flowers. In
grasses Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns ...
, each floret (flower) is enclosed in a pair of papery bracts, called the lemma (lower bract) and palea (upper bract), while each spikelet (group of florets) has a further pair of bracts at its base called glumes. These bracts form the chaff removed from cereal grain during
threshing Threshing, or thrashing, is the process of loosening the edible part of grain (or other crop) from the straw to which it is attached. It is the step in grain preparation after reaping. Threshing does not remove the bran from the grain. History ...
and
winnowing Winnowing is a process by which chaff is separated from grain. It can also be used to remove pests from stored grain. Winnowing usually follows threshing in grain preparation. In its simplest form, it involves throwing the mixture into the ...
. Bats may detect acoustic signals from dish-shaped bracts such as those of '' Marcgravia evenia''. A prophyll is a leaf-like structure, such as a bracteole, subtending (extending under) a single flower or pedicel. The term can also mean the lower bract on a peduncle. The frequently showy pair of bracts of ''
Euphorbia ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
'' species in subgenus ''Lacanthis'' are the cyathophylls. Bracts subtend the cone scales in the seed cones of many conifers, and in some cases, such as '' Pseudotsuga'', they extend beyond the cone scales. File:Castilleja levisecta bracts.jpg, Bright yellow bracts of ''
Castilleja levisecta ''Castilleja levisecta'' is a rare species of flowering plant in the family Orobanchaceae known by the common name golden paintbrush, or golden Indian paintbrush, listed under the Endangered Species Act in 1997. It is native to British Columbia a ...
'' are more tightly pressed against the stem than bracts of other ''
Castilleja ''Castilleja'', commonly known as paintbrush, Indian paintbrush, or prairie-fire, is a genus of about 200 species of annual and perennial herbaceous plants native to the west of the Americas from Alaska south to the Andes, northern Asia, and on ...
'' species. File:Bougainvillea glabra.JPG, Bracts of ''
Bougainvillea glabra ''Bougainvillea glabra'', the lesser bougainvillea or paperflower, is the most common species of bougainvillea used for bonsai. The epithet 'glabra' comes from Latin and means "bald". Description It is an evergreen, climbing shrub with thick ...
'', differ in colour from the non-bract leaves, and attract pollinators. File:Banana bract.jpg, Bracts along a banana flower stalk surround the rows of flowers File:E milii vulcanii ies.jpg, ''
Euphorbia milii ''Euphorbia milii'', the crown of thorns, Christ plant, or Christ thorn, is a species of flowering plant in the spurge family Euphorbiaceae, native to Madagascar. The species name commemorates Baron Milius, once Governor of Réunion, who introdu ...
'' var. ''vulcanii'' cyathia bearing a pair of pinkish cyathophylls. File:Ananas bracteatus var tricolor.jpg, Colourful bracts of ''
Ananas bracteatus The pineapple (''Ananas comosus'') is a tropical plant with an edible fruit; it is the most economically significant plant in the family Bromeliaceae. The pineapple is indigenous to South America, where it has been cultivated for many centurie ...
''. File:Erigeron peregrinus 6287.JPG, The many green involucral bracts of ''
Erigeron peregrinus ''Erigeron peregrinus'' is a North American species of flowering plants in the family Asteraceae known by the common name wandering fleabane. ''Erigeron peregrinus'' is native to northwestern North America from Alaska to Oregon. Some populations ...
'' are linear, loose, taper to a point, about the same length, and help to distinguish this species. File:Blooming Dogwood.jpg,
Dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shru ...
species '' Cornus florida'' inflorescence showing four large white bracts and central flower cluster. File:2013-05-10 08 26 08 Closeup of pink dogwoods at the Brendan T. Byrne State Forest headquarters.jpg, '' Cornus florida'' dogwood
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
with pink bracts surrounding small flower cluster. File:豔紅苞鳳梨 20190712192436.jpg, The red bracts of ''
Aechmea bracteata ''Aechmea bracteata'' is a plant species in the genus ''Aechmea''. This species is native to Central America, Mexico, Colombia, and Venezuela; it is also reportedly naturalized in the Bahamas The Bahamas (), officially the Commonwealth ...
''. File:女王鬱金 20190715203108 01.jpg, The pink bracts of '' Curcuma petiolata''. File:火鶴 20190729181532.jpg, The pink bract of a new cultivar: '' Anthurium andraeanum'' 'Pink Lady'. File:豔山姜花序總苞 20190508131956.jpg, The inflorescence of '' Alpinia zerumbet'' protected by two bracts before flower bloom. File:地湧金蓮 Musella lasiocarpa 20201028130818 01.jpg, The golden bracts of ''
Musella lasiocarpa ''Musella lasiocarpa'' ( syn. ''Musa lasiocarpa''), commonly known as Chinese dwarf banana, golden lotus banana or Chinese yellow banana, is the sole species in the genus ''Musella''. It is thus a close relative of bananas, and also a member of ...
''. File:Hydrangea involucrata (9344379310).jpg, Large bracts of '' Hydrangea involucrata'' surround tiny flowers and demonstrate the "lacecap" style
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
that has made various ''
Hydrangea ''Hydrangea'', () commonly named the hortensia, is a genus of over 75 species of flowering plants native to Asia and the Americas. By far the greatest species diversity is in eastern Asia, notably China, Korea, and Japan. Most are shrubs tall, ...
'' species and cultivars popular with gardeners. File:Viburnum opulus RF.jpg, Like many ''
Hydrangea ''Hydrangea'', () commonly named the hortensia, is a genus of over 75 species of flowering plants native to Asia and the Americas. By far the greatest species diversity is in eastern Asia, notably China, Korea, and Japan. Most are shrubs tall, ...
'' species (
Hydrangeaceae Hydrangeaceae is a family of flowering plants in the order Cornales, with a wide distribution in Asia and North America, and locally in southeastern Europe. Description The genera are characterised by leaves in opposite pairs (rarely whorled ...
family), ''
Viburnum opulus ''Viburnum opulus'', the guelder-rose or guelder rose () is a species of flowering plant in the family Adoxaceae (formerly Caprifoliaceae) native to Europe, northern Africa and central Asia. Description ''Viburnum opulus'' is a deciduous s ...
'' (
Adoxaceae Adoxaceae, commonly known as moschatel family, is a small family of flowering plants in the order Dipsacales, now consisting of five genera and about 150–200 species. They are characterised by opposite toothed leaves, small five- or, more rare ...
family) has bracts that surround tiny flowers, creating a "lacecap" style
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
.


Bracteole

A small bract is called a bracteole or bractlet. Technically this is any bract that arises ''on'' a pedicel instead of subtending it.


Involucral bracts

Bracts that appear in a whorl subtending an
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
are collectively called an involucre. An involucre is a common feature beneath the inflorescences of many
Apiaceae Apiaceae or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus '' Apium'' and commonly known as the celery, carrot or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering plan ...
,
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
,
Dipsacaceae The Dipsacaceae have been recognized as a family (the teasel family) of the order Dipsacales containing 350 species of perennial or biennial herbs and shrubs in eleven genera. The species are currently placed in the family Caprifoliaceae. Native ...
and
Polygonaceae The Polygonaceae are a family of flowering plants known informally as the knotweed family or smartweed—buckwheat family in the United States. The name is based on the genus '' Polygonum'', and was first used by Antoine Laurent de Jussieu in 1 ...
. Each flower in an inflorescence may have its own whorl of bracts, in this case called an involucel. In this case they may be called ''chaff'', ''paleas'', or ''receptacular bracts'' and are usually minute scales or bristles. Many asteraceous plants have bracts at the base of each inflorescence. The term involucre is also used for a highly conspicuous bract or bract pair at the base of an inflorescence. In the family
Betulaceae Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the ...
, notably in the genera ''
Carpinus Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbea ...
'' and ''
Corylus The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ...
'', the involucre is a leafy structure that protects the developing nuts. Beggar-tick ('' Bidens comosa'') has narrow involucral bracts surrounding each inflorescence, each of which also has a single bract below it. There is then a pair of leafy bracts on the main stem and below those a pair of leaves.


Epicalyx

An epicalyx, which forms an additional whorl around the calyx of a single flower, is a modification of bracteoles In other words, the epicalyx is a ''group of bracts resembling a calyx'' or ''bracteoles forming a whorl outer to the calyx''. It is a calyx-like extra whorl of floral appendages. Each individual segment of the epicalyx is called an episepal because they resemble the sepals in them. They are present in the hibiscus family,
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familiar ...
. '' Fragaria'' (strawberries) may or may not have an epicalyx.


Spathe

A ''spathe'' is a large bract or pair of bracts forming a sheath to enclose the flower cluster of such plants as
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...
s,
arum ''Arum'' is a genus of flowering plants in the family Araceae, native to Europe, northern Africa, and western and central Asia, with the highest species diversity in the Mediterranean region. Frequently called arum lilies, they are not closely ...
s, irises,Richard Lynch
crocus ''Crocus'' (; plural: crocuses or croci) is a genus of seasonal flowering plants in the family Iridaceae (iris family) comprising about 100 species of perennials growing from corms. They are low growing plants, whose flower stems remain under ...
es,Alex Casha and
dayflower ''Commelina'' is a genus of approximately 170 species commonly called dayflowers due to the short lives of their flowers. They are less often known as widow's tears. It is by far the largest genus of its family, Commelinaceae. The Swedish taxonom ...
s ('' Commelina''). '' Zephyranthes tubispatha'' in the Amaryllidaceae derives its specific name from its tubular spathe. In many arums (family
Araceae The Araceae are a family of monocotyledonous flowering plants in which flowers are borne on a type of inflorescence called a spadix. The spadix is usually accompanied by, and sometimes partially enclosed in, a spathe (or leaf-like bract). A ...
), the spathe is petal-like, attracting pollinators to the flowers arranged on a type of spike called a spadix.


References

{{Botany Plant morphology Leaves