Spanish imperial eagle
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The Spanish imperial eagle (''Aquila adalberti''), also known as the Spanish eagle or Adalbert's eagle, is a species of
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, j ...
native to the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
. The binomial commemorates Prince Adalbert of Bavaria. Due to its distinct “epaulettes”, old literature often referred to this species as the white-shouldered eagle. Formerly, the Iberian imperial eagle was considered to be a subspecies of the
eastern imperial eagle The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South ...
, but is now widely recognised as a separate species due to differences in
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
,
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
, and molecular characteristics.


Description

This is a large raptor and quite large eagle, broadly similar in size to its cousin, the
eastern imperial eagle The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South ...
, which is found in a considerably different distributional range. Compared to sympatric largish booted eagles, it is somewhat smaller than the
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds of ...
and somewhat larger than the
Bonelli's eagle The Bonelli's eagle (''Aquila fasciata'') is a large bird of prey. The common name of the bird commemorates the Italian ornithologist and collector Franco Andrea Bonelli. Bonelli is credited with gathering the type specimen, most likely from an ...
. Spanish imperial eagle can weigh from . The average weight of males in a sample of 10 was while that of 17 females was found to be . Meanwhile, another sample of 10 unsexed adults weighed an average of . Thus, the Spanish imperial eagle weighs about 10% more on average than the eastern imperial eagle and rivals the considerably longer-winged and longer-tailed
wedge-tailed eagle The wedge-tailed eagle (''Aquila audax'') is the largest bird of prey in the continent of Australia. It is also found in southern New Guinea to the north and is distributed as far south as the state of Tasmania. Adults of this species have lon ...
as the third heaviest member of the '' Aquila'' genus behind the golden and
Verreaux's eagle Verreaux's eagle (''Aquila verreauxii'') is a large, mostly African, bird of prey. It is also called the black eagle, especially in southern Africa, not to be confused with the Indian black eagle (''Ictinaetus malayensis''), which lives far to t ...
s. This species has a total length of and a wingspan of .European Raptor Conservation European This is a national bird of
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

Spanish Imperial Eagle, Aquila adalberti.
' Accessed 17 May 2011
Thiollay, J. M., Del Hoyo, J., Elliott, A., & Sargatal, J. (1994). ''Handbook of the Birds of the World, vol. 2., New World Vultures to Guineafowl''. Lynx Editions.González, L. M. (2016). ''Águila imperial ibérica – Aquila adalberti''. En: ''Enciclopedia Virtual de los Vertebrados Españoles''. Salvador, A., Morales, M. B. (Eds.). Museo Nacional de Ciencias Naturales, Madrid. A typical wingspan for a male is reportedly about while for a female may be about . The adult resembles the eastern imperial eagle and can superficially suggest the golden eagle (especially when distantly seen), but is overall a darker color than either, a rich blackish-brown which extends all the way from the throat down to the belly. Like the eastern imperial, the adult has a broad distinctive white band on the shoulder and leading edge of the wing, which is even more pronounced in the Spanish than in the eastern species, and a much paler tawny color on the nape and crown, unlike the golden-yellow color on a similar area in the golden eagle. The juvenile Spanish imperial eagle is very different from adults and other large raptors in this range, being overall a uniform pale straw-sandy colour, contrasting with broad black bands on both the upper and lower sides of the wings. Smaller than the relatively small race of golden eagle found in the Iberian peninsula, it is somewhat slighter and slenderer in appearance compared to the more powerful golden species, with a relatively longer neck, and generally much flatter wing profile in flight than the upturned dihedral typical of a golden eagle.Ferguson, L. J., & Christie, D. A. (2001). ''Raptors of the World''. London (UK): Christopher Helm.


Range

The species occurs in central and south-west
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
and adjacent areas of
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
, in the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
. Its stronghold is in the dehesa woodlands of central and south-west Spain, such as in
Extremadura Extremadura (; ext, Estremaúra; pt, Estremadura; Fala: ''Extremaúra'') is an autonomous community of Spain. Its capital city is Mérida, and its largest city is Badajoz. Located in the central-western part of the Iberian Peninsula, it ...
, Ciudad Real and areas in the north of
Huelva Huelva (, ) is a city in southwestern Spain, the capital of the province of Huelva in the autonomous community of Andalusia. It is between two short rias though has an outlying spur including nature reserve on the Gulf of Cádiz coast. The ria ...
and Seville's Sierra Norte. The Spanish imperial eagle is a resident species, unlike the partially migratory eastern imperial eagle. Stable occurrence in
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
is disputed but immature birds during the dispersion period regularly visit Morocco. Rising numbers of vagrant birds born in Spain and then electrocuted in Morocco have been noted; some areas used by the species in Morocco could be becoming sort of a "drain" in terms of the species recovery and this is due to the fact that the country stands in a similar situation as Spain was in the early 1980s when it comes to insulation of transmission towers.Las águilas imperiales vuelven a Marruecos y mueren electrocutadas, Ciencia, EL PAÍS
/ref> Vagrant birds have even reached Mauritania and
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
. North of its natural range, vagrants have reached as far as the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
in one rare occasion.


Ecology

Nesting habitat is usually dry, mature woodlands, which they utilize for nesting and seclusion, but nests are most often fairly close to shrubby openings and wetland areas where prey is more likely to be concentrated. A shy species toward man, they normally nest only where human disturbance is quite low. Like most raptors, they are highly territorial and tend to maintain a stable home range. Spanish imperial eagles nest from February to April. The nesting pair will construct a nest of as much as across at first build, which will increase in time, especially in mature ''
Quercus suber ''Quercus suber'', commonly called the cork oak, is a medium-sized, evergreen oak tree in the section ''Quercus'' sect. ''Cerris''. It is the primary source of cork for wine bottle stoppers and other uses, such as cork flooring and as the core ...
'' or pine trees. Clutch size is usually two to three eggs, with an incubation period of about 43 days, but on average about 1.23-1.4 fledglings are produced per nest. Nestling mortality is usually due to human disturbance and destruction and nest collapses, secondarily due to predation and
siblicide Siblicide (attributed by behavioural ecologist Doug Mock to Barbara M. Braun) is the killing of an infant individual by its close relatives (full or half siblings). It may occur directly between siblings or be mediated by the parents, and is dri ...
. Fledging is reached at 63–77 days of age but juveniles can linger for an extremely long period, to at least 160 days after fledging. It feeds mainly on
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has b ...
s, which comprised about 58% of this species' diet before
myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus '' Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North A ...
and
rabbit haemorrhagic disease Rabbit hemorrhagic disease (RHD), also known as viral hemorrhagic disease (VHD), is a highly infectious and lethal form of viral hepatitis that affects European rabbits. Some viral strains also affect hares and cottontail rabbits. Mortality rate ...
greatly reduced the rabbit's native Iberian population. As rabbit population crashed they've been recorded feeding on a wide range of
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () (chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with c ...
s with varied success depending upon prey populations and may become semi-specialized hunters of
water bird A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from seabi ...
s especially
Eurasian coot The Eurasian coot (''Fulica atra''), also known as the common coot, or Australian coot, is a member of the rail and crake bird family, the Rallidae. It is found in Europe, Asia, Australia, New Zealand and parts of North Africa. It has a slaty-bla ...
s,
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s and
geese A goose ( : geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera '' Anser'' (the grey geese and white geese) and ''Branta'' (the black geese). Some other birds, mostly related to the she ...
, also taking some numbers of partridges, pigeons and crows and any other bird they happen to encounter that is vulnerable to ambush. More than 60 bird species are known in be included in their prey spectrum. Several mammals may too be taken occasionally including various rodents, Granada hare, hares, mustelids, hedgehogs and even other large predators such as red foxes or—rarely, since they are not typically present in the eagle's habitat—domestic cats and small dogs. Rarely, reptiles or even fish may also be preyed upon. The largest prey taken by this species may easily exceed , such as foxes, greylag geese or white storks, but mean prey mass is relatively low, especially in areas with fewer rabbits. One study reported mean prey mass as locally, though average prey size has also been reported more highly. The Spanish imperial eagle is one of several rabbit-favoring birds of prey in Spain along with the similarly specialized Iberian lynx. This species is largely segregated by habitat from other eagles that specialize on rabbits here to lessen direct competition, as the imperial eagle favors woods, whereas the golden and Bonelli's eagles tend to dwell in much rockier areas. However, Spanish imperial eagles frequently quarrel over food with various raptors, even much larger Old World vultures, vultures, and the raptors may at times try to kill the young of one another. In one case, in protection their own nest, an adult Spanish imperial eagle even killed a cinereous vulture, the largest accipitrid in the world. Healthy, free-flying Spanish imperial eagles are apex predators, being mostly free of natural predators themselves but they do sometimes kill each other in conflicts and rarely interspecies conflicts may too be fatal. When protected from human persecution and far from threats such as powerlines, adult mortality can be as low as 3–5.4% annually.


Conservation

The species is classified as Vulnerable species, Vulnerable by IUCN. Threats include loss of habitat, human encroachment, collisions with Transmission tower, pylons (at some point in the early 1980s, powerlines were responsible for 80% of deaths among birds in their first year of life) and illegal poisoning. There has also been a decline in the species' main prey: rabbits have been kept at bay or even declined in some of the areas where the eagle is or could be present as a result of
myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus '' Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North A ...
and, most recently,
rabbit haemorrhagic disease Rabbit hemorrhagic disease (RHD), also known as viral hemorrhagic disease (VHD), is a highly infectious and lethal form of viral hepatitis that affects European rabbits. Some viral strains also affect hares and cottontail rabbits. Mortality rate ...
. By the 1960s it had become a critically endangered species, with only 30 pairs remaining, all located in Spain. Following conservation efforts, recovery began in the 1980s at a rate of five new breeding pairs per year up to 1994. Imperial eagles were nearly wiped out. In 2011, the species's global population had increased to 324 pairs, with 318 pairs in Spain. The species recolonised Portugal in 2003, after an absence of breeding activity for over 20 years, and has been slowly increasing since, with six breeding pairs located in 2011 and nine located in 2012. The population in Spain showed an average annual increase of c. 7% between 1990 and 2011. These positive trends are largely attributed to mitigation measures to reduce mortality associated with powerlines, supplementary feeding, reparation of nests, reintroductions and decreases in the disturbance of breeding birds, although some of the observed increases may be due to more thorough searches within its range.Tintó, A., Real, J., & Mañosa, S. (2010). ''Predicting and correcting electrocution of birds in Mediterranean areas''. Journal of Wildlife Management, 74(8), 1852-1862.


Notes


References

* * * Ferguson-Lees, James & Christie, David A. (2001): ''Raptors of the World''. Houghton Mifflin, Boston. . * Ferrer, Miguel (2001): ''The Spanish Imperial Eagle''. Lynx Edicions, Barcelona. . * * * * Schuhmacher, Eugen (1973): ''Europe's Paradises'' *


External links


BirdLife Species Factsheet
{{Authority control Aquila (genus), Spanish imperial eagle Birds of Southern Europe Fauna of the Iberian Peninsula Birds of North Africa Birds described in 1861, Spanish imperial eagle Taxa named by Christian Ludwig Brehm, Spanish imperial eagle