Sordes
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''Sordes'' was a small
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
from the late
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
( Oxfordian/
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
)
Karabastau Svita The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...
of
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
. This genus was named in 1971 by
Aleksandr Grigorevich Sharov Aleksandr Grigorevich Sharov (А.Г. Шаров, 1922–1973) was a Soviet palaeoentomologist, paleontologist and expert on Pterosauria. He graduated from Moscow State University. In 1951 he defended Candidate of Science dissertation on the embryo ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
is ''Sordes pilosus''. The genus name is
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
for "filth" or "scum"; but Sharov translates it as "", which means "devil" or "evil spirit", so the intended translation is "hairy devil" (the specific name is Latin for "hairy"; despite ''sordes'' being feminine, it has not yet been emended to ''pilosa'').


Discovery

''Sordes'' is based on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
PIN 2585/3, which consists of a crushed relatively complete skeleton on a slab. It was found in the 1960s at the foothills of the
Karatau Karatau ( kz, Қаратау, ''Qaratau'') is an administrative center in the Talas region located in the Jambyl Region in the south of Kazakhstan. It has an estimated population of roughly 30,000 in 1999. , the population is now 27,667. The la ...
in
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
. Sharov had already referred a
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...
or second specimen: PIN 2470/1, again a fairly complete skeleton on a slab. By 2003 another six specimens had been discovered.


Description

''Sordes'' had a 0.63 m (2 ft) wingspan. The wings were relatively short. ''Sordes'' had, according to Sharov and Unwin, wing membranes attached to the legs and a membrane between the legs. It had a short neck. It had a long tail, accounting for over half its length, with at the end an elongated vane.


Skull and dentition

It had a slender, not round, head with moderately long, pointed jaws. The skull was about 8 cm (3.2 in) long. Unlike many pterosaurs, it had no head crest. The teeth in the frontal half of the jaws are large and pointed to facilitate prey capture. The teeth beyond these in the rear half of the jaw are much smaller and more numerous than those at the front, suggesting that they were more for crushing. Together these two types of teeth indicate specialisation for prey that was difficult to catch yet required some effort to eat. Likely contenders are invertebrates with tougher exoskeletons, or amphibians that were slippery to catch and then required some crunching before they could be swallowed.


Pycnofibers

The
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
shows remains of the soft parts, such as membranes and hair-like filaments. This was the first unequivocal proof that pterosaurs had a layer of hair-like filaments covering their bodies, later named pycnofibres. The pycnofibres served as insulation, an indication the group was
warm-blooded Warm-blooded is an informal term referring to animal species which can maintain a body temperature higher than their environment. In particular, homeothermic species maintain a stable body temperature by regulating metabolic processes. The on ...
, and provided a streamlined flight profile. The pycnofibres are present in two main types: longer at the extreme part of the wing membrane and shorter near the body. In the 1990s,
David Unwin David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...
argued that both types were essentially not hairs but reinforcing fibres of the flight membranes.Elgin, R.A., Hone, D.W.E., and Frey, E. 2011. The extent of the pterosaur flight membrane. Acta Palaeontologica Polonica 56 (1): 99–111

/ref> Later he emphasized that "hair" in the form of pycnofibres was indeed present on the body, after the find of new specimens clearly showing this.


Classification

''Sordes'' has been assigned to the family
Rhamphorhynchidae Rhamphorhynchidae is a group of early pterosaurs named after ''Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study o ...
. These were among the earliest of the pterosaurs, evolving in the late
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest per ...
and surviving to the late Jurassic. According to Unwin, within Rhamphorhynchidae ''Sordes'' belonged to the
Scaphognathinae Rhamphorhynchidae is a group of early pterosaurs named after ''Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study o ...
. Other researchers however, such as
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
and
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
, have produced
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analyses showing that ''Sordes'' was much more basal, and not a rhamphorhynchid.


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inval ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...


References


External links


Artistic reconstruction of a ''Sordes pilosus'' at the Fossilsmith StudiosA fossil of a ''Sordes'' at paleo.ru
{{Portalbar, Paleontology Late Jurassic pterosaurs of Asia Novialoids Taxa named by Aleksandr Grigorevich Sharov Fossil taxa described in 1971