Sanderling
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The sanderling (''Calidris alba'') is a small wading bird. The name derives from
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th ...
''sand-yrðling'', "sand-ploughman". The genus name is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
''kalidris'' or ''skalidris'', a term used by
Aristotle Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of ...
for some grey-coloured waterside birds. The specific, ''alba'', is
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
for "white". It is a circumpolar
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
breeder, and is a long-distance migrant, wintering south to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
, South
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
. It is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. It is somewhat unlike other
sandpiper Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil ...
s in appearance, which has led to the suggestion that it should be placed into a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
genus ''Crocethia''. A more recent review (Thomas ''et al.'', 2004) indicates, however, that the sanderling is a fairly typical "
stint A stint is one of several very small waders in the paraphyletic ''"Calidris"'' assemblage – often separated in ''Erolia'' – which in North America are known as peeps. They are scolopacid waders much similar in ecomorphology to the ...
" or small sandpiper and should be separated from the large knots with its closest relatives in a distinct genus. This bird is similar in size to a
dunlin The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brow ...
, but stouter, with a thick bill. It shows a strong white wingbar in flight, and runs along the sandy beaches it prefers with a characteristic "bicycling" action of its legs, stopping frequently to pick small food items. It eats small
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...
s and other small
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s. In spring, birds migrating north from South America consume large numbers of
horseshoe crab Horseshoe crabs are marine and brackish water arthropods of the family Limulidae and the only living members of the order Xiphosura. Despite their name, they are not true crabs or crustaceans: they are chelicerates, most closely related to ar ...
eggs in the
Delaware Bay Delaware Bay is the estuary outlet of the Delaware River on the northeast seaboard of the United States. It is approximately in area, the bay's freshwater mixes for many miles with the saltwater of the Atlantic Ocean. The bay is bordered inland ...
area. In spring, the birds arrive on the High Arctic breeding grounds (see map), where they lay 3–4 eggs in a ground scrape. On the nesting grounds, these birds mainly eat
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s and some plant material. The sanderling was described by the German naturalist
Peter Simon Pallas Peter Simon Pallas FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son of Professor of Surgery ...
in 1764 and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''Trynga alba''. The sanderling is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (
AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Conventio ...
) applies.


Description

The sanderling is a small plump sandpiper, in length. Its weight ranges from . The winter
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
is very pale, almost white apart from a dark shoulder patch. This is the source of the specific name, ''alba'', which is the Latin for "white". Later in the summer, the face and throat become brick-red. The juvenile bird is spangled black and white, and shows much more contrast than the adult. If its size is misjudged, a sanderling in breeding plumage can be mistaken for some varieties of
stint A stint is one of several very small waders in the paraphyletic ''"Calidris"'' assemblage – often separated in ''Erolia'' – which in North America are known as peeps. They are scolopacid waders much similar in ecomorphology to the ...
, or a sanderling in winter plumage can be mistaken for a
dunlin The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brow ...
or
red knot The red knot or just knot (''Calidris canutus'') is a medium-sized shorebird which breeds in tundra and the Arctic Cordillera in the far north of Canada, Europe, and Russia. It is a large member of the '' Calidris'' sandpipers, second only to the ...
. It can be told from other small wading birds, given good views, by its lack of a hind toe. Its behavior is also distinctive.


Distribution, habitat and migration

The sanderling breeds in the High Arctic areas of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
. In North America, it breeds in the
Canadian Arctic Archipelago The Arctic Archipelago, also known as the Canadian Arctic Archipelago, is an archipelago lying to the north of the Canadian continental mainland, excluding Greenland (an autonomous territory of Denmark). Situated in the northern extremity of ...
,
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
,
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
(and to a lesser extent
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
). In
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
, it breeds in
Spitsbergen Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern Nor ...
and areas of northern Russia from the
Taymyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrat ...
to the
New Siberian Islands The New Siberian Islands ( rus, Новосиби́рские Oстрова, r=Novosibirskiye Ostrova; sah, Саҥа Сибиир Aрыылара, translit=Saña Sibiir Arıılara) are an archipelago in the Extreme North of Russia, to the north ...
. In the northern winter, it has a nearly
cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The extr ...
across the world's marine coasts. It is a complete migrant, travelling between from its breeding grounds to its wintering sites. Birds that travel further also arrive later and leave sooner. Most adults leave the breeding grounds in July and early August, whereas juvenile birds leave in late August and early September. The northward migration begins in March at the southern end of their winter distribution. The breeding habitat of the sanderling is coastal
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
north of July isotherm. The species typically chooses nesting sites on dry stony areas near wet areas, from above sea level to . During the winter and its migration, it is most commonly found on coastal sandy
beach A beach is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock, such as sand, gravel, shingle, pebbles, etc., or biological sources, such as mollusc s ...
es, but also occurs on tidal sand flats, mud flats and the shores of lakes and rivers. More infrequently, it may occur on rocky shores.


Subspecies

The sanderling consists of two subspecies: *''C. a. alba'', (
Pallas Pallas may refer to: Astronomy * 2 Pallas asteroid ** Pallas family, a group of asteroids that includes 2 Pallas * Pallas (crater), a crater on Earth's moon Mythology * Pallas (Giant), a son of Uranus and Gaia, killed and flayed by Athena * Pa ...
, 1764)
, breeds on
Ellesmere Island Ellesmere Island ( iu, script=Latn, Umingmak Nuna, lit=land of muskoxen; french: île d'Ellesmere) is Canada's northernmost and third largest island, and the tenth largest in the world. It comprises an area of , slightly smaller than Great Br ...
, north & east
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
,
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group rang ...
,
Franz Josef Land , native_name = , image_name = Map of Franz Josef Land-en.svg , image_caption = Map of Franz Josef Land , image_size = , map_image = Franz Josef Land location-en.svg , map_caption = Location of Franz Josef ...
and the
Taymyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrat ...
*''C. a. rubida'', ( Gmelin, 1789), breeds in northeast
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
,
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
and northern
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...


Behaviour


Feeding behaviour

Sanderlings feed on invertebrate
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
buried in the sand in the upper
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
. In North America, this diet largely consists of the
isopods Isopoda is an order of crustaceans that includes woodlice and their relatives. Isopods live in the sea, in fresh water, or on land. All have rigid, segmented exoskeletons, two pairs of antennae, seven pairs of jointed limbs on the thorax, and ...
'' Excirolana linguifrons'', '' Excirolana kincaidii'', and the mole crab, ''
Emerita analoga ''Emerita analoga'', the Pacific sand crab or Pacific mole crab, is a species of small, sand-burrowing decapod crustacean found living in the sand along the temperate western coasts of North and South America. It is found on exposed sandy beache ...
''. When the tide is out, these
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s live in burrows some way beneath the surface. When the tide comes in, they move into the upper layers of sand and feed on the
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
and
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts comm ...
that washes over them with each wave. They then burrow rapidly down again as the water retreats. They leave no marks on the surface, so the sanderlings hunt for them by plunging their beaks into the sand at random, consuming whatever they find. Their bills can penetrate only and as the water swirls around and retreats, the sand is softer; this makes it easier for the birds' beaks to penetrate further. In the spring, when much breeding activity is taking place in the
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning " ...
community, there may be as many as 4000 invertebrates per square metre, but their average size is smaller than later in the year. The birds appear to rush madly around at the edge of the surf, but in reality they are maximising their chances of catching as many prey animals as possible when they are at their most vulnerable near the surface.


Breeding behaviour

At breeding time sanderlings are
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
, with the male aggressively defending its territory. They may either form monogamous pairs or
polyandrous Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wive ...
(one female and two male) pairings.


Gallery

File:Sanderling Dan Pancamo.jpg, Sanderling at
High Island, Texas High Island is an unincorporated community located in the Bolivar Peninsula census-designated place, Galveston County, Texas, United States. The community is located in the extreme eastern part of the county on Bolivar Peninsula, less than one mi ...
File:SANDERLING.jpg, A Sanderling in
Cagayan de Oro Cagayan ( ), officially the Province of Cagayan ( ilo, Probinsia ti Cagayan; ibg, Provinsiya na Cagayan; itv, Provinsiya ya Cagayan; fil, Lalawigan ng Cagayan), is a province in the Philippines located in the Cagayan Valley region, covering ...
,
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
File:SanderlingsRunningFromWavesCropped.jpg, A flock displaying their distinctive behavior of running with the ebb and flow of waves (while feeding). Willapa Bay, near
Tokeland, Washington Tokeland is a census-designated place (CDP) in Pacific County, Washington, United States. The population was 151 at the 2010 census, a sharp decrease from the 194 at the 2000 census. The town was named after Chief Toke, an Indian chief of th ...
. File:OneLeggedSanderling.jpg, Resting at the
Monterey Bay Aquarium Monterey Bay Aquarium is a nonprofit public aquarium in Monterey, California. Known for its regional focus on the marine habitats of Monterey Bay, it was the first to exhibit a living kelp forest when it opened in October 1984. Its biolo ...
aviary An aviary is a large enclosure for confining birds, although bats may also be considered for display. Unlike birdcages, aviaries allow birds a larger living space where they can fly; hence, aviaries are also sometimes known as flight cages. Avi ...
. File:Sanderling-floreana.jpg, Floreana Island, Galapagos Islands File:Sanderling (with tag).jpg, A banded bird File:Calidris-alba-001.jpg, Running File:Sanderling from the Crossley ID Guide Britain and Ireland.jpg, ID composite File:Sanderling-11MAY2017.JPG, Sanderling in Monterey, California File:Sanderling (Calidris alba).jpg, Amrum.jpg, Sanderling on the island of Amrum, Schleswig-Holstein File:Sanderlings (72649).jpg, Sanderlings feeding in Quogue, New York


Notes


Footnotes


References

*


External links


Sanderling Species text in The Atlas of Southern African Birds
*

— Cornell Lab of Ornithology

— USGS Patuxent Bird Identification InfoCenter * * * * * * * * {{Authority control Calidris Erolia Sandpipers Birds of the Arctic Birds of the Dominican Republic Birds of the Caribbean Cosmopolitan birds Birds described in 1764 Taxa named by Peter Simon Pallas Holarctic birds