Sand goanna
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The sand goanna (''Varanus gouldii'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of large
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
n monitor
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...
, also known as Gould's monitor, sand monitor, or racehorse goanna.


Taxonomy

John Edward Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used f ...
described the species in 1838 as ''Hydrosaurus gouldii'', noting the source of the type specimen as "New Holland" and distinguishing the new varanid by "two yellow streaks on the sides of the neck" and small flat scales at the orbits. An earlier description, ''Tupinambis endrachtensis'' Péron, F. 1807, was determined as likely to refer to this animal, but the epithet ''gouldii'' was conserved and a new specimen designated as the type. This neotype was obtained in 1997 at the near coastal Western Australian suburb of Karrakatta, and placed with the
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum ...
. The decision of a nomenclatural commission (
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
) was to issue an opinion suppressing the earlier names, ''Tupinambis endrachtensis'' and ''Hydrosaurus ocellarius'' Blyth, 1868, that were unsatisfactory to some who had commented on the case, but provided taxonomic certainty for future revisions of the associated taxa. The specific name is assumed to be a Latinised form of an associate of the describing author, the
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist. He published a number of monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould, and several other artists, including Edward Lear, ...
, who was actively assembling specimens of fauna from Australia but is not thought to have any direct connection to this species. In some Aboriginal languages, the sand goanna is called ''bungarra'', a term also commonly used by non-Aboriginal people in
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to t ...
. In
Pitjantjatjara The Pitjantjatjara (; or ) are an Aboriginal people of the Central Australian desert near Uluru. They are closely related to the Yankunytjatjara and Ngaanyatjarra and their languages are, to a large extent, mutually intelligible (all are vari ...
and other central Australian languages it is called "Tingka". Two subspecies are recognised, *'' Varanus gouldii gouldii'' – Gould's goanna *'' Varanus gouldii flavirufus'' – desert sand monitor ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atte ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
other than ''Varanus''.


Description

A species of '' Varanus'', lizards known as monitors and goannas, that is found in a variety of habitat. Due to the taxonomic uncertainty during the twentieth century the species form and behaviour has included taxa later recognised as distinct species, this includes '' V. rosenbergi'', formerly treated as a subspecies and later elevated, and '' V. panoptes'', described as a new species in 1980 and resolved as a legitimate publication in 2000.


Biology

Gould's monitor is a terrestrial reptile that excavates large
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of s ...
s for shelter. Rock escarpments and tree hollows are also suitable dwellings. It inhabits a vast range throughout Australia, reaching an average length of and weighing as much as . It can be found in northern and eastern Australia, where they inhabit open
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s and
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s. ''V. g. flavirufus'', a slightly smaller
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
, resides in Australia's interior. In some places, however, the
ranges In the Hebrew Bible and in the Old Testament, the word ranges has two very different meanings. Leviticus In Leviticus 11:35, ranges probably means a cooking furnace for two or more pots, as the Hebrew word here is in the dual number; or perhaps ...
of Gould's monitor, ''V. g. flavirufus'' and the Argus monitor overlap. The similarities between the species and their close proximity frequently cause confusion. The sand monitor is a relentless forager. It is diurnal, meaning most of its activities take place during the day. Anything smaller than itself will be eagerly devoured. The diets of hatchlings and juveniles often consist mostly of
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s and small
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...
s, but generally varies more with age. Adult monitors will
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
on
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus'' ...
, other small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s, small
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s,
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s, large insects, small lizard species,
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
s, amphibians, eggs, and carrion. They are seemingly immune to snake venom and will kill and eat even the most venomous species such as the inland taipan. The sand goanna does consume smaller species of monitors: Ackies, rock monitors and other dwarf species are often found and eaten. It is common to see a Gould's monitor disturbing rock piles in an attempt to flush out any '' Odatria''. It lays its eggs in
termite mounds Mound-building termites are a group of termite species that live in mounds. These termites live in Africa, Australia and South America. The mounds sometimes have a diameter of . Most of the mounds are in well-drained areas. Termite mounds usual ...
to protect them from the harsh
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
climate. Like all goannas and
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
s the sand monitor has a forked tongue, with which it "tastes" the air. Scents are picked up on each side of the fork which, when retracted, brushes over an area in the mouth called the
Jacobson's organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. T ...
, which identifies on which side the scent is stronger. Upon identifying this difference in scent strength the sand monitor can alter its direction accordingly. A 2021 study had shown that during the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
,
Komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a member of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo, Rinca, Flores, and Gili Motang. It is the largest extant ...
s had hybridized with the ancestors of the sand goanna, thus providing evidence that the Komodo dragon had once inhabited Australia.


Gallery

File:Sand Goanna near Broken Hill. 1976.jpg, Sand goanna near Broken Hill File:Sand monitor walking tracks in sand.jpg, Sand monitor walking tracks in sand. File:Bangara at Gidgee.jpg, Bungarra at the Gidgee Gold Mine. File:Varanus gouldii at Sunrise Dam Gold Mine.jpg, Bungarra at the Sunrise Dam Gold Mine. File:Varanus gouldii - Chace ranges.JPG, Chace ranges, South Australia File:Sand Goanna on road (15734829198).jpg, Mungo National Park, NSW, Australia File:Varanus gouldii camera lure.JPG, Sand goanna at a camera lure File:Sand goanna.jpg, Sand goanna File:Varanus-gouldii.jpg, Northern Territory


Citations


General sources

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External links


Photo of ''Varanus gouldii'' at AROD.com.au

Photo of ''Varanus gouldii'' at Pbase.com
{{Taxonbar, from=Q671373 Varanus Reptiles described in 1838 Taxa named by John Edward Gray Goanna, sand Reptiles of Queensland Reptiles of New South Wales Monitor lizards of Australia