Salix caprea
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''Salix caprea'', known as goat willow,
pussy willow Pussy willow is a name given to many of the smaller species of the genus ''Salix'' (willows and sallows) when their furry catkins are young in early spring. These species include (among many others): *Goat willow or goat sallow ('' Salix caprea ...
or great sallow, is a common species of
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...
native to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and western and central
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook 4. .


Description

It is a
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...
or small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
, reaching a height of , rarely to 13 m. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are 3–12 cm long and from 2–8 cm wide, broader than most other willows. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are soft silky, and silvery 3-7-cm-long
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
s are produced in early spring before the new leaves appear; the male and female catkins are on different plants (
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
). The male catkins mature yellow at
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
release, the female catkins mature pale green. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a small capsule 5–10 mm long containing numerous minute
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s embedded in fine, cottony hairs. The seeds are very small (about 0.2 mm) with the fine hairs aiding dispersal; they require bare
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
to
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins. . The two
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
are: *''S. c.'' var. ''caprea'' - lowland regions throughout the range, leaves thinly hairy above, densely hairy below, 5–12 cm long, stipules persistent until autumn *''S. c.'' var. ''sphacelata'' (Sm.) Wahlenb. (syn. ''S. caprea'' var. ''coaetanea'' Hartm.; ''S. coaetanea'' (Hartm.) Floderus) - high altitudes in the mountains of central and northern Europe (Alps, Carpathians, Scotland, Scandinavia), leaves densely silky-hairy on both sides, 3–7 cm long, stipules early deciduous. In western Norway there are trees of S. c. var 'Sphacelata' that are 18-20 meters tall and have mostly single trunks.


Name

The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''caprea'' means "goat". This, and the common name goat willow, probably derive from the first known illustration of the species in
Hieronymus Bock Hieronymus Bock ( Latinised Hieronymus Tragus; c. 1498 – 21 February 1554) was a German botanist, physician, and Lutheran minister who began the transition from medieval botany to the modern scientific worldview by arranging plants by their re ...
's 1546 '' Herbal'', where the plant is shown being browsed by a goat. The species was historically also widely used as a browse for goats, to which Bock's illustration may refer.Bean, W. J. (1980). ''Trees and Shrubs Hardy in the British Isles''. .


Ecology

''S. caprea'' occurs both in wet/damp environments, such as riverbanks and lake shores, and in drier sites, wherever bare soil becomes available due to ground disturbance. Hybrids with several other willow species are common, notably with '' Salix cinerea'' (''S. × reichardtii''), '' Salix aurita'' (''S. × multinervis''), '' Salix viminalis'' (''S. × smithiana''), and '' Salix purpurea'' (''S. × sordida''). Populations of ''S. caprea'' often show hybrid introgression. Unlike almost all other willows, pure specimens do not take root readily from cuttings; if a willow resembling the species does root easily, it is probably a hybrid with another species of willow. The leaves are used as a food resource by several species of
Lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described speci ...
, and are also commonly eaten by browsing
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s. Willows are very susceptible to gall inducers, and the midge '' Rhabdophaga rosaria'' forms the camellia gall on ''S. caprea''.Gall Inducers


Cultivation and uses

A small number of
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have been selected for garden use. The most common is ''S. caprea'' 'Kilmarnock', discovered by James Smith, with stiffly pendulous shoots forming a mop-head; it is a male clone. A similar female clone is ''S. caprea'' 'Weeping Sally'. As they do not form a leader, they are
grafted Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
on erect stems of other willows; the height of these cultivars is determined by the height at which the graft is made. Plants can also be grown from greenwood cuttings, which make attractive creeping mounds. Hardwood cuttings are often difficult to root. Both
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
and salicin can be extracted from goat willow bark. The tree is not considered a good source of
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
, as its wood is both brittle and known to crackle violently if burned. As with the closely related ''
Salix discolor ''Salix discolor'', the American pussy willow or glaucous willow, is a species of willow native to North America, one of two species commonly called pussy willow. It is native to the vast reaches of Alaska as well as the northern forests and wet ...
'' (American pussy willow), it is also often grown for cut flowers. See
Pussy willow Pussy willow is a name given to many of the smaller species of the genus ''Salix'' (willows and sallows) when their furry catkins are young in early spring. These species include (among many others): *Goat willow or goat sallow ('' Salix caprea ...
for further cultural information, which applies to both species. In Scandinavia it has been fairly common to make willow flutes from goat willow cuttings. In
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwee ...
,
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Cr ...
, north of
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,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inva ...
, the just-opened catkins are used like the olive branches on
Palm Sunday Palm Sunday is a Christian moveable feast that falls on the Sunday before Easter. The feast commemorates Christ's triumphal entry into Jerusalem, an event mentioned in each of the four canonical Gospels. Palm Sunday marks the first day of Hol ...
.


References


External links


''Salix caprea''
- genetic conservation units
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote '' in situ'' an ...
(EUFORGEN) * * {{Taxonbar, from=Q159565 caprea Flora of Ukraine Plants described in 1753 Taxa named by Carl Linnaeus