''Raeticodactylus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of non-

pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...

from the late

Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...

-early

Rhaetian The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest a ...

-age

Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. ...

lower

Kössen Formation The Kössen Formation is a Late Triassic (Rhaetian-age) geological formation in the Northern Calcareous Alps of Austria and Germany, in the Tiroler-Lech Nature Park. During the Late Triassic, the area now occupied by the Northern Calcareous Alps w ...

of the central Austroalpine of

Grisons The Grisons () or Graubünden,Names include: *german: (Kanton) Graubünden ; * Romansh: ** rm, label= Sursilvan, (Cantun) Grischun ** rm, label= Vallader, (Chantun) Grischun ** rm, label= Puter, (Chantun) Grischun ** rm, label= Surmiran, (Ca ...

Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...

. It is known from

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

BNM 14524, a single disarticulated partial skeleton including an almost complete skull, found in August 2005. This genus was named and described in 2008 by its discoverer Rico Stecher; the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

is ''Raeticodactylus filisurensis''. The specific name refers to

Filisur Filisur is an Alpine village and former municipality in the Albula Region in the canton of Graubünden in Switzerland. The village sits on a hillside with a view to the west where the two rivers Albula/Alvra from the Albula Pass and Landwasser ...

. Online First

Description

''Raeticodactylus'' had a tall thin bony crest running along the midline of the front of the upper jaw, and a keel on the lower jaw; however, it does not seem to be closely related to ''

Austriadactylus ''Austriadactylus'' is a genus of "rhamphorhynchoid" pterosaur. The fossil remains were unearthed in Late Triassic (middle Norian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: ...

'', the only other crested Triassic pterosaur named by the time ''Raeticodactylus'' was described. The teeth at the front of the upper jaw, in the

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

e, were fanglike, whereas the teeth in the upper cheeks (the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

e) had three, four, or five cusps, similar to those of ''

Eudimorphodon ''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (m ...

''. ''Raeticodactylus'' had a wingspan of about 135

centimeters 330px, Different lengths as in respect to the Electromagnetic spectrum, measured by the Metre and its deriveds scales. The Microwave are in-between 1 meter to 1 millimeter. A centimetre (international spelling) or centimeter (American spellin ...

(53 in), and may have been a piscivore, potentially feeding by skimming the water. However, skim-feeding has since been disproven in pterosaurs, and the related '' Caviramus'' appears to have been an omnivore.

Classification

A 2009 study by Fabio Dalla Vecchia, which analyzed the relationships within the family

Campylognathoididae Novialoidea (meaning "new wings") is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous (early Toarcian to late Maastrichtian ageBarrett, P. M., Butler, R. J., Edwards, N. P ...

, found that ''Raeticodactylus'' is very closely related to '' Caviramus''. ''Caviramus'' is known only from a partial jaw, exhibiting the same type of teeth and unique cavities in the jaw bone, and both specimens come from the same time and place (the Alplihorn Member of the Swiss Kössen Formation). Dalla Vecchia concluded that the two belong to the same genus, and possibly the same species, if the subtle differences (such as size and the presence of a crest in the ''Raeticodactylus'' specimen) are not due to sex or age. A study by

Brian Andres Brian (sometimes spelled Bryan in English) is a male given name of Irish and Breton origin, as well as a surname of Occitan origin. It is common in the English-speaking world. It is possible that the name is derived from an Old Celtic word me ...

''et al.'' in 2014 concluded that ''Raeticodactylus'' was the

sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of ''Caviramus'' within the family

Raeticodactylidae Raeticodactylidae is a family of eudimorphodontoid eopterosaurian pterosaurs that lived in Switzerland during the Late Triassic. The family includes '' Caviramus'', and the type genus '' Raeticodactylus'', which are both known from the Kössen F ...

. The following phylogenetic analysis follows the topology of Andres ''et al.'' (2014). In 2020 however, a study upheld by Matthew G. Baron about early pterosaur interrelationships found ''Raeticodactylus'' to group with ''

Carniadactylus ''Carniadactylus'' is a genus of pterosaur which existed in Europe during the Late Triassic period (late Carnian or early Norian, about 228 million years agoBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribu ...

'', ''Caviramus'', and the clade Austriadraconidae, which in turn were within a larger clade called Caviramidae.Matthew G. Baron (2020). "Testing pterosaur ingroup relationships through broader sampling of avemetatarsalian taxa and characters and a range of phylogenetic analysis techniques". PeerJ. 8: e9604. doi:10.7717/peerj.9604. PMC 7512134. .

References

{{Taxonbar, from=Q1405497 Pterosaurs Late Triassic pterosaurs of Europe Fossil taxa described in 2008 Bergün Filisur

Description

Classification

See also

References